Land snail diversity in the monsoon tropics of Northern Australia: revision of the genus Exiligada Iredale, 1939 (Mollusca: Pulmonata: Camaenidae), with description of 13 new species Author Criscione, Francesco Author Law, Margot Louisa Author Köhler, Frank text Zoological Journal of the Linnean Society 2012 Zool. J. Linn. Soc. 2012-11-26 166 4 689 722 http://dx.doi.org/10.1111/j.1096-3642.2012.00863.x journal article 10.1111/j.1096-3642.2012.00863.x 0024-4082 5409181 AC086999-76A8-4F93-9A51-5B5B213A50E6 EXILIGADA QUALIS IREDALE, 1939 Figure 19. Penial anatomy of E. nodulicauda sp. nov. (holotype WAM S83173). Scale bar = 5 mm. ( FIGS 4E , 6E , 23A–C , 24 ) Exiligada qualis Iredale, 1939: 69 , plate V, figure 2. Exiligada negriensis – Solem, 1984: 673–677 , figure 174 (d–f) (in part). Figure 20. SEM micrographs of Exiligada shells.A-C, E. pallida sp. nov. (paratype WAM S49150 ). A, Apical view showing protoconch and first whorl. B, Close-up from above of sculpture across entire shell. C, Microsculpture on teleoconch whorl. D-F, E. punctata sp. nov. (paratype WAMS49139). D, Apical view showing protoconch and first whorl. E, Close-up from above of sculpture across entire shell. F, Microsculpture on teleoconch whorl. Scale bars: A-B, D-E = 1 mm, C, F = 0.5 mm. Material examined: AUSTRALIA , NT , VRD , holotype from 25 miles north of Ord River Station , Negri outstation (coll. R. Helms , 1896), dry ( AM C.64866), 43 paratypes (same data as for holotype) ( AM C.64915); 35 km north of Nicholson H /S, Duncan Rd, limestone cliffs, 500 m west of road, under rocks at dripline of cave overhang, 17°42 ′ 45 ′′ S , 128°50 ′ 10 ′′ E (coll. R. Crookshanks , 28.i.2005 ), three dry, one wet ( AMS C.443304) . Duncan Rd , old Ord River homestead, Forrest Ck , 70 km north of Nicholson homestead, under limestone rocks and concrete of old structures, 17°23 ′ 56 ′′ S , 128°52 ′ 01 ′′ E (coll. R. Crookshanks , 29.i.2005 ), one wet ( AM C.475758) . Description: Shell ( Figs 4E , 6E , 23A–C ). Moderately large, moderately elevated, moderately thin; teleoconch with short regularly alternate microdepressions. Shell background colour yellowish horn fading to whitish towards shell base, with few conspicuous to moderately faint continuous brown-reddish spiral bands; maculations absent. Genitalia ( Fig. 24 ). Epiphallus as long as penis. Vas deferens entering penial sheath from halfway up. Penis six times longer than wide, not coiled inside sheath, with inner wall supporting a large apical stimulatory pilaster and (several) basal pilasters. Vagina two thirds of penis and free oviduct, bursa copulatrix with no well-differentiated end. Remarks: The type locality is identical to that of E. negriensis and as such also restricted to an area close to the Negri River between the point where the Negri River flows into the Ord River in WA ( 17°04 ′ 11 ′′ S , 128°53 ′ 10 ′′ E ) and the Duncan Highway bridge over the Negri River near the WA / NT border relative to width (pl/pw) and epiphallus length relative to penis (el/pl) average. Inner penial wall sculpture similar to those of E. brabyi , to be distinguished by posterior pilaster being continuous. Vagina length relative to penis (vl/pl) average and vagina length relative to free oviduct (vl/ol) half the average.