<document id="C21414BF9BB441A9EF5B5C45DEAB9BA5" ID-DOI="10.1649/0010-065X-72.3.433" ID-ISSN="1938-4394" ID-Zenodo-Dep="5383463" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630296642708" checkinUser="diego" docAuthor="Gillett, Conrad P. D. T. &amp; Barr, Iain" docDate="2018" docId="03E32325FF803362858A3CB5FBDAFCC4" docLanguage="en" docName="0010-065X-72.3.433.pdf" docOrigin="The Coleopterists Bulletin 72 (3)" docSource="http://dx.doi.org/10.1649/0010-065x-72.3.433" docStyle="DocumentStyle:F824235CA89CB83023C7D5789122F870.3:TheColeopBull.2017-.journal_article.1cover" docStyleId="F824235CA89CB83023C7D5789122F870" docStyleName="TheColeopBull.2017-.journal_article.1cover" docStyleVersion="3" docTitle="Heliocopris pirmal" docType="treatment" docVersion="3" lastPageNumber="436" masterDocId="FFDA5B5DFF84336687633D48FFE8FFB5" masterDocTitle="New Country Records of Scarab Dung Beetles (Coleoptera: Scarabaeidae: Scarabaeinae) in Swaziland and Zambia" masterLastPageNumber="438" masterPageNumber="433" pageNumber="436" updateTime="1699219109578" updateUser="plazi">
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<mods:title id="4037E96C175ABABFFB6C399E0D0FB56D">New Country Records of Scarab Dung Beetles (Coleoptera: Scarabaeidae: Scarabaeinae) in Swaziland and Zambia</mods:title>
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<taxonomicName id="4C4AE9B0FF803362858A3CB5FBB1FDA6" ID-CoL="3KB4N" authority="(Fabricius, 1798)" baseAuthorityName="Fabricius" baseAuthorityYear="1798" box="[745,1113,509,531]" class="Insecta" family="Scarabaeidae" genus="Heliocopris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="436" phylum="Arthropoda" rank="species" species="pirmal">
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<emphasis id="B93E4E21FF803362858A3CB5FC4CFDA7" bold="true" box="[745,932,509,530]" italics="true" pageId="4" pageNumber="436">Heliocopris pirmal</emphasis>
(Fabricius, 1798)
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Previously recorded from 
<collectingCountry id="F35DD2A3FF80336284B53F60FBF6FD88" box="[982,1054,552,573]" name="Angola" pageId="4" pageNumber="436">Angola</collectingCountry>
, 
<collectingCountry id="F35DD2A3FF80336283533F60FB78FD88" box="[1072,1168,552,573]" name="Botswana" pageId="4" pageNumber="436">Botswana</collectingCountry>
, 
<collectingCountry id="F35DD2A3FF80336285CE3F0BFCC5FDED" box="[685,813,579,600]" name="Mozambique" pageId="4" pageNumber="436">Mozambique</collectingCountry>
, 
<collectingCountry id="F35DD2A3FF80336284563F0BFC61FDED" box="[821,905,579,600]" name="Namibia" pageId="4" pageNumber="436">Namibia</collectingCountry>
, 
<collectingCountry id="F35DD2A3FF80336284F33F0BFBE3FDED" box="[912,1035,579,600]" name="South Africa" pageId="4" pageNumber="436">South Africa</collectingCountry>
, 
<collectingCountry id="F35DD2A3FF80336283713F0BFB82FDED" box="[1042,1130,579,600]" name="Tanzania" pageId="4" pageNumber="436">Tanzania</collectingCountry>
, and 
<collectingCountry id="F35DD2A3FF80336285CE3F15FCFAFDC7" box="[685,786,605,626]" name="Zimbabwe" pageId="4" pageNumber="436">Zimbabwe</collectingCountry>
(
<bibRefCitation id="EFDBEFC2FF80336284423F15FC4FFDC7" author="Ferreira, M. C." box="[801,935,605,626]" pageId="4" pageNumber="436" pagination="5 - 1088" refId="ref3778" refString="Ferreira, M. C. 1972. Os escarab'ideos de Africa (sul do Saara). I. Revista de Entomologia de Mocambique 11: 5 - 1088." type="journal article" year="1972">Ferreira 1972</bibRefCitation>
; 
<bibRefCitation id="EFDBEFC2FF80336284D13F15FB9FFDC7" author="Pokorny, S." box="[946,1143,605,627]" pageId="4" pageNumber="436" refId="ref4119" refString="Pokorny, S., J. Z'idek, and K. Werner. 2009. Giant Dung Beetles of the Genus Heliocopris (Scarabaeidae). Taita Publishers, Hradec Kralove, Czech Republic." type="book" year="2009">
Pokorný 
<emphasis id="B93E4E21FF80336283693F16FBD4FDC6" box="[1034,1084,605,627]" italics="true" pageId="4" pageNumber="436">et al.</emphasis>
2009
</bibRefCitation>
).
</paragraph>
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<emphasis id="B93E4E21FF80336285A63F30FC7AFD38" bold="true" box="[709,914,632,653]" pageId="4" pageNumber="436">New country record</emphasis>
<materialsCitation id="3B22986EFF80336284F93F30FC24FCA7" collectingDate="2016-03-25" country="Swaziland" location="One" municipality="Simunye" pageId="4" pageNumber="436" specimenCount="1">
for 
<collectingCountry id="F35DD2A3FF80336284DD3F30FBCAFD38" box="[958,1058,632,653]" name="Swaziland" pageId="4" pageNumber="436">Swaziland</collectingCountry>
: 
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specimen, 
<collectingMunicipality id="6B910849FF80336285823FDBFCDFFD1D" box="[737,823,659,680]" pageId="4" pageNumber="436">Simunye</collectingMunicipality>
, 
<date id="FFF4B4F3FF803362845C3FDBFC38FD1D" box="[831,976,659,680]" pageId="4" pageNumber="436" value="2016-03-25">
<collectingDate id="EFB04D1BFF803362845C3FDBFC38FD1D" box="[831,976,659,680]" pageId="4" pageNumber="436" value="2016-03-25">25 March 2016</collectingDate>
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. A female was found crawling around an outdoor shopping center in company with specimens of 
<taxonomicName id="4C4AE9B0FF80336284A23F8FFCCAFD42" authority="Bates. They" authorityName="Bates. They" class="Insecta" family="Scarabaeidae" genus="Heliocopris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="436" phylum="Arthropoda" rank="species" species="andersoni">
<emphasis id="B93E4E21FF80336284A23F8FFB7BFD68" box="[961,1171,711,733]" italics="true" pageId="4" pageNumber="436">Heliocopris andersoni</emphasis>
Bates. They
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apparently had been attracted to the lights there the previous night
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.
</paragraph>
<paragraph id="8BF59233FF803362858F3E6FFBDAFCC4" blockId="4.[707,1150,807,881]" pageId="4" pageNumber="436">
Tribe Onthophagini Burmeister, 1846 Genus 
<taxonomicName id="4C4AE9B0FF803362846A3E09FBDAFCC4" authority="Roggero, Dierkens, Barbero, and Palestrini, 2017" authorityName="Roggero, Dierkens, Barbero, and Palestrini" authorityYear="2017" class="Insecta" family="Scarabaeidae" genus="Hamonthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="436" phylum="Arthropoda" rank="genus">
<emphasis id="B93E4E21FF803362846A3E09FC5EFCE2" box="[777,950,833,855]" italics="true" pageId="4" pageNumber="436">Hamonthophagus</emphasis>
Roggero, Dierkens, Barbero, and Palestrini, 2017
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