An overview of the extant genera and subgenera of the order Scolopendromorpha (Chilopoda): a new identification key and updated diagnoses Author Schileyko, Arkady A. schileyko1965@gmail.com Author Vahtera, Varpu varpu.vahtera@gmail.com Author Edgecombe, Gregory D. 0000-0002-9591-8011 schileyko1965@gmail.com text Zootaxa 2020 2020-08-10 4825 1 1 64 journal article 8703 10.11646/zootaxa.4825.1.1 5ab5f5c8-481e-4d1a-8643-21e72c367278 1175-5326 4402145 F230F199-1C94-4E2E-9CE4-5F56212C015F Family Scolopendridae Leach, 1814 Diagnosis. Four ocelli ( Fig. 52 ) form a “rhomboid cluster” (the only blind exceptions—the genus Tonkinodentus and a clade of Western Australian Cormocephalus species; see below). Labrum with a single median tooth (fig. 7 of Waldock & Edgecombe 2012 ). Pretarsus of maxillae 2 not pectinate, robust and straight ( Figs 59 , 74 , 80 ), accompanied by 2 (more rarely by 0 or 1) accessory spine(s). Anterior margin of forcipular coxosternite ( Figs 40, 42 , 50 , 86 ) with tooth-plates (except for the monotypic genus Edentistoma ; Fig. 114 ). Forcipular trochantero-prefemur practically always with well-developed process, the latter may be simple or furnished by 1–3 lateral tubercles. Sternites in most genera with longitudinal paramedian sutures. 21 LBS (21, 23, 39, 43 in Scolopendropsis only); spiracles on macrosegments, LBS 7 with or without spiracles. Ultimate LBS ( Fig. 38 ) generally considerably shorter than the penultimate one (except for Scolopendropsis , Fig. 45 ). Tarsus of locomotory legs with two articles; legs with or without 1 or 2 tibial spurs and 1 tarsal spur. Coxopleuron usually with process ( Fig. 49 ). Ultimate legs in most genera and species of “common” shape, rarely “pincer-shaped” or “leaf-shaped” (sensu Schileyko 2009 ). Prefemur of the ultimate legs without spinous processes but generally with varying number (2–20) of strongly chitinized spines, that are usually small ( Figs 44 , 49 , 66 , 70 , 83 ) but rarely much enlarged, for example in Kanparka (= Scolopendra ) ( Fig. 56 ) or a few forms of Ethmostigmus Pocock, 1898 ( Fig. 105 , figs 36, 39 in Schileyko & Stoev 2016 ). Edgecombe & Bonato (2011: 395) also wrote: “Labral bristle field completely covering distal sclerotisation of epipharynx (except Notiasemus ; [no data on Tonkinodentus ]); elongate, figure eight shaped groups of two smooth depressions surrounding each sensillum on clypeal part or epipharynx. Tufts of bristles on lateral flaps of hypopharynx form a continuous field with identical bristles medially… Poison calyx extending at least as far as proximal part of forcipular trochantero-prefemur… Spermatophore with a ventral invagination.” Number of subtaxa. 2 subfamilies, 19 genera (“More than 400 species in 21 genera” sensu Edgecombe & Bonato 2011: 397 ). Sexual dimorphism. Present rarely. Range. All tropical, subtropical and warm temperate regions. Remarks. Treated as a family in Edgecombe & Bonato (2011: 395) , Kronmüller (2012: 19) , Edgecombe et al. (2012: 770) , Vahtera et al. (2012a: 4 , 2012b: 235 , 2013: 578 ), Schileyko (2014: 174) , Schileyko & Stoev (2016: 252) , Schileyko (2018: 69), Schileyko & Solovyeva (2019: 138) .