An overview of the extant genera and subgenera of the order Scolopendromorpha (Chilopoda): a new identification key and updated diagnoses
Author
Schileyko, Arkady A.
schileyko1965@gmail.com
Author
Vahtera, Varpu
varpu.vahtera@gmail.com
Author
Edgecombe, Gregory D.
0000-0002-9591-8011
schileyko1965@gmail.com
text
Zootaxa
2020
2020-08-10
4825
1
1
64
journal article
8703
10.11646/zootaxa.4825.1.1
5ab5f5c8-481e-4d1a-8643-21e72c367278
1175-5326
4402145
F230F199-1C94-4E2E-9CE4-5F56212C015F
(!)
Rhysida
H.C. Wood, 1862
Figs 106–109
Type
species.
Branchiostoma lithobioides
Newport
, 1845
(by subsequent designation of
Attems 1930
)
.
Diagnosis.
Median tooth of labrum well developed. Forcipular tooth-plates present, trochantero-prefemur with well-developed process (
Fig. 106
). Tergites never with longitudinal keels (except for median one in a few species). Sternites with incomplete paramedian sutures (in some species much shortened, sometimes also with median and lateral depressions of various sizes/shapes. LBS 7 with spiracles, the latter with atrium (
Fig. 108
). Legs with tarsal spur(s), legs 1 without prefemoral spur. Coxopleural process (
Fig. 109
) ranging from short to very long and much enlarged (figs 4F and 9F in
Siriwut
et al
. 2018
, respectively), with spines (at least with apical ones). Prefemur of the ultimate leg with spines (
Fig. 107
), more rarely (for example in
R. celeris
) without them; pronounced corner spine absent (in some species 1(2) spines at its place,
Fig. 107
). Ultimate pretarsus well-developed, with accessory spines.
Number of species.
36 (
Joshi
et al.
2020
).
Sexual dimorphism.
Unknown.
Remarks.
Treated as a genus in
Edgecombe & Bonato (2011: 402)
,
Vahtera
et al.
(2012a: 7
,
2012b: 238
,
2013: 578
), Chagas-Jr (2013: 17),
Schileyko (2014: 182)
,
Schileyko & Stoev (2016: 255)
,
Siriwut
et al.
(2018: 1005)
;
Joshi
et al
. (2020)
; the latter work is the most recent account on this genus (see also Remarks to
Alluropus
below). In 2013 Chagas-Jr synonymized seven species of
Rhysida
(namely
R. caripensis
,
R. guayanica
,
R. maritima
,
R. monaguensis
,
R. neoesparanta
,
R. porlamarensis
and
R. sucupaensis
) described by
González-Sponga (2002)
to
R. celeris
. Five new species were described from
India
by
Joshi
et al.
(2020)
.
We re-investigated a dozen specimens of
R. celeris
from
Peru
(Rc 6685) and
Brazil
(Rc 7272),
R. immarginata
(Porat, 1876)
from
Papua New Guinea
(Rc 7091),
R. longipes
(Newport, 1845)
from
Cambodia
(Rc 7003) and
Peru
(Rc 6683),
R. lithobioides
(Newport, 1845)
from Sumatra (Rc 7232). All the studied specimens showed a total absence of both a corner spine of the ultimate legs (
Fig. 107
) and a prefemoral spur on legs 1.