A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae) Author Park, J. - S. Louisiana State Arthropod Museum, Department of Entomology, LSB 404 Louisiana State University Agric Author Carlton, C. E. Louisiana State Arthropod Museum, Department of Entomology, LSB 404 Louisiana State University Agric text The Coleopterists Bulletin 2014 mo 13 2014-12-01 68 1 156 journal article 3079 10.1649/072.068.0mo4.1 62322993-ca80-4b6c-ac95-861cbb236abd 1938-4394 4907735 Sagola insignis Broun, 1893 ( Figs. 68a , 69a , 70a , 71 ) Sagola insignis Broun, 1893b: 1049 . Raffray 1904: 497 , 1911: 5 , 1924: 233 ; Hudson 1923: 365 , 1934: 183 ; Newton and Thayer 2005b; Nomura and Leschen 2006: 242. Sagola sobrina Broun, 1893b: 1050 . Hudson 1923: 365 , 1934: 183 ; Raffray 1924: 233 ; Newton and Thayer 2005b; Nomura and Leschen 2006: 243. New Synonymy . Type Material. Holotype . NEW ZEALAND : Northland : 1♂ ( BMNH ), glued on rectangular card, “Type” [red label, printed]; “1875.” [white label, handwritten]; “Mokohinau” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola insignis ” [white label, handwritten] . Syntypes of Sagola sobrina : NEW ZEALAND : Northland : 1♀ ( BMNH ), glued on rectangular card, “Type” [red label, printed]; “1877.” [white label, handwritten]; “Mokohinau” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola sobrina ” [white label, handwritten]. 1♀ ( BMNH ), glued on rectangular card, “1877.” [white label, handwritten]; “Mokohinau I” [white label, handwritten]; “ New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]. 1♀ ( BMNH ), glued on rectangular card, “1877. ♀ ” [white label, handwritten]; “Mokohinau I” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed] . Additional Material ( n = 35: 12 males ; 23 females ). NEW ZEALAND : Auckland : 2♂♂ , Lynfield WB , 13 VI 1980 , G. Kuschel; 1♀ , Lynfield, 28 VIII 1976 , G. Kuschel, litter; 1♂ , Manurewa , Murphy’ s Bush , 2 V 1981, G. Kuschel, rotten wood; 1♂ , 20–40m , Grafton Gully , 4 III 2001 , S.E. Thorpe, bush litter; Coromandel : 5♂♂ 9♀♀ , The Alderman I , Ruamahuaiti I , 40m , 6–11 XII 1994 , J.W. Early , R .F. Gilbert , yellow pan trap ; 1♂ 1♀ , Little Barrier I, 3 I 1952 ; 1♀ , Little Barrier I, 2 I 1952 ; Northland : 4♂♂ 1♀ , Mokohinau I , Trig I , 27 II 1978 , G. Kuschel, litter; 2♀♀ , Poor Knights I , Aorangi , ridge to Oneho Hill , 17 XI 1981 , J.C. Watt, litter; 2♀♀ , Poor Knights I , Tawhiti Rahi , 4 XII 1980 , G. Kuschel, litter; 1♀ , Poor Knights I , Aorangi , Puweto Valley , 14 XI 1981 , J.C. Watt, litter; 1♀ , Hen & Chickens I , Lady Alice I, I 1982 , L. Roberts, litter; 1♀ , Poor Knights I , Aorangi , Puweto Valley , 9–17 XI 1981 , J.C. Watt, pit trap; 1♀ , Poor Knights I , Aorangi , Crater Bay , 9–17 XI 1981 , J.C. Watt, pit trap. Diagnosis. This species can be distinguished from the other species of the insignis speciesgroup by the following combination of characters: body length 2.6–2.9 mm ; temporal lobe of male head distinctly projecting, exceeding eye, with concave ventral surface; ventral surface of male head with transverse process behind mouthparts with three patches of dense setae medially; male elytra rectangular, female elytra subquadrate; shape of antennomeres and genitalia unique to species. Fig. 68. Habitus of Sagola species in the insignis species-group, dorsal views. a) S. insignis , b) S. hectorii , c) S. bipunctata , d) S. angulifera , e) S. eminens , f) S. robustula , g) S. pertinax , h) S. monticola , i) S. laticeps , j) S. castanea , k) S. sheldoni , l) S. mercuryensis , m) S. cuvierensis , n) S. buckleyi , o) S. brumfieldi , p) S. otteai , q) S. boudreauxae , r) S. taegyui , s) S. hanae , t) S. gyudongi , u) S. egmontensis , v) S. boonei , w) S. stewartensis , x) S. jeongnamae . Scale bars = 1 mm. Redescription. Length 2.6–2.9 mm . Body brown, antennae, legs, maxillary palpi, and elytra paler ( Fig. 68a ). Head: Male head triangular, widest across temples ( Fig. 69a ). Female head bluntly triangular, widest across eyes. Antennomere 1 approximately 1.5 times longer than wide, 2 longer than wide, 3 subquadrate, 4–6 longer than wide, 7–10 subquadrate. Frontal sulcus deep, reaching end of eye. Anterior frontal fovea round, partially covered by frontal rostrum, posterior frontal fovea round. Eye prominent, approximately one-third length of temple. Temporal lobe of male head distinctly projecting, exceeding eye with concave ventral surface ( Fig. 69a ). Ventral surface of male head with transverse process behind mouthparts with 3 patches of dense setae medially ( Fig. 69a ). Thorax: Prosternum as long as wide, widest at one-third length. Male elytra rectangular ( Fig. 68a ), female elytra subquadrate. Male meso-and metathorax trapezoidal, longer than wide, female as long as wide. Abdomen: Male tergite IV with pair of small, round patches of microtrichia, patches absent in female. Aedeagus: Median lobe of genitalia broad with 3 round depressions apically ( Fig. 70a ). Phallobase of median lobe symmetrical and rounded ( Fig. 70a ). Parameres symmetrical and slender, setose from midpoints to apices ( Fig. 70a ). Fig. 69. Ventral views of male heads of Sagola species in the insignis species-group. a) Sagola insignis , b) S. hectorii , c) S. bipunctata , d) S. angulifera , e) S. eminens , f) S. robustula , g) S. pertinax , h) S. monticola , i) S. laticeps , j) S. castanea , k) S. sheldoni , l) S. mercuryensis , m) S. cuvierensis , n) S. buckleyi , o) S. brumfieldi , p) S. otteai , q) S. boudreauxae , r) S. taegyui , s) S. hanae , t) S. gyudongi , u) S. egmontensis , v) S. boonei , w) S. stewartensis , x) S. jeongnamae . Scale bars = 0.1 mm. Fig. 70. Sagola species in the insignis species-group. Aedeagi, dorsal views: a) S. insignis , b) S. hectorii , c) S. bipunctata , d) S. angulifera , e) S. eminens , f) S. robustula , g) S. pertinax , h) S. monticola , i) S. laticeps , j) S. castanea , k) S. sheldoni , l) S. mercuryensis , m) S. cuvierensis , n) S. buckleyi , o) S. brumfieldi , p) S. otteai , q) S. boudreauxae , r) S. taegyui , s) S. hanae , t) S. gyudongi , u) S. egmontensis , v) S. boonei , w) S. stewartensis , x) S. jeongnamae . Male heads, dorsal surface: y) S. bipunctata , z) S. eminen . Scale bars = 0.1 mm. Type Locality. Mokohinou Island , New Zealand . Distribution. Auckland , Coromandel, Northland ( Fig. 71 : black circles). Habitat. Specimens of this species were collected using pitfall and yellow pan traps or by sifting leaf litter. Comments. Specimens of S. insignis can be distinguished from those of other species by the shape of the ventral surface of the male head and the antennomeres. The type specimens of S. sobrina share these diagnostic characters, and additional specimens have been collected at or near the type locality. For these reasons, we have placed S. sobrina in synonymy with S. insignis .