Haematophagous biting midges of the extant genus Culicoides Latreille (Diptera: Ceratopogonidae) evolved during the mid-Cretaceous Author Szadziewski, Ryszard Author Dominiak, Patrycja Author Sontag, Elżbieta Author Krzemiński, Wiesław Author Wang, Bo Author Szwedo, Jacek text Zootaxa 2019 2019-10-24 4688 4 535 548 journal article 25149 10.11646/zootaxa.4688.4.5 c704c5d3-8ef6-48b6-a346-efe5b5e1c032 1175-5326 3517710 F412A472-E501-494B-9BF9-CAF253E9BD92 Culicoides burmiticus Szadziewski & Dominiak, sp. nov. Figs 4 A–B urn:lsid:zoobank.org:act: 8F5AD5B1-78A5-491A-8036-FD4EC46EAD2A Diagnosis. This new species is distinguished from other Cretaceous Culicoides by the following female characters: antennal flagellum with cylindrical flagellomeres, flagellomeres 2–8 much shorter than elongate 9–13, only flagellomere 1 with single sensillum coeloconicum; wing membrane except basal radial cell with macrotrichia; palpal segment 3 stout with distinct, deep sensory pit; proboscis moderately short. Male unkown. Description. Female ( Fig. 4A ). Very well preserved, all structures present and well visible. Head-body length 1.03 mm . Eyes narrowly separated. Antennal flagellum ( Fig. 4B ) length 0.54 mm ; flagellomere 1 with single sensillum coeloconicum, flagellomeres 2–13 cylindrical without sensilla coecolonica; antennal ratio 1.44. Palpus ( Fig. 4B ) 5-segmented; segment 3 stout 0.045 mm long, with deep sensory pit. Wing ( Fig. 4A ) length measured from basal arculus 0.70 mm ; costal ratio 0.69; wing membrane except basal radial cell with macrotrichia. Legs slender, hind femur most massive; hind tibial comb with 3 spines; tarsomeres 4 cylindrical; tarsal ratios similar on all legs, TR (1) 2.2, TR (2) 2.5, TR (3) 2.3; claws small, equal-size. Male. Unknown. Material examined. Holotype female, BUB 1745, Burmese amber, Hukawng Valley, Myanmar . Syninclusions. Collembola 2, Acari 1, Empididae 1 female , Ceratopogonidae : Leptoconops 3 females . Deposited in Museum of Natural History, PAS Kraków. Etymology. This new species name refers to Burma ( Myanmar ), the country of origin of the amber in Southeast Asia.