Hieracium moravense (Asteraceae), a new hawkweed from Albania
Author
Gottschlich, Günter
0000-0003-0677-6478
Hermann-Kurz-Strasse 35, 72074 Tübingen, Germany & ggtuebingen @ yahoo. com; https: // orcid. org / 0000 - 0003 - 0677 - 6478
ggtuebingen@yahoo.com
Author
Selvi, Federico
0000-0002-3820-125X
Department of Agriculture, Food, Environment and Forestry, Laboratories of Botany, University of Firenze, Piazzale delle Cascine 28, I- 50144, Firenze, Italy & National Biodiversity Future Center, Palermo, Italy & federico. selvi @ unifi. it; https: // orcid. org / 0000 - 0002 - 3820 - 125 X
federico.selvi@unifi.it
text
Phytotaxa
2023
2023-04-13
592
1
73
78
http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN
journal article
249067
10.11646/phytotaxa.592.1.7
bc5d13a8-232c-40ed-af6a-76b16867ea96
1179-3163
7835716
Hieracium moravense
Gottschl. & Selvi
,
spec. nov.
(„waldsteinii – bifidum“) (figs. 1+2)
FIGURE 1.
Hieracium moravense
,
spec. nov.
Left: holotype (FI), right: isotype (Hb. Gottschlich-81174).
FIGURE 2.
Hieracium moravense
, single capitulum.
Type
:—
ALBANIA
.
Korcë
,
Mt. Moravë
, gola rocciosa che parte
da Drenovë
, rocce e rupi ultramafiche (serpentino),
40.5780°N
,
20.7966°E
,
980–1100 m
,
14.07.2022
,
F
. Selvi,
A
. Coppi,
I
. Colzi,
E
. Bianchi s. n
. (
holotype
:
FI068417
; isotype:
Hb. Gottschlich
81174)
.
Diagnosis
:—
Planta inter
Hieracium waldsteinii
subsp
. plumulosum
et
H. bifidum
intermedia
, ab
Hieracio waldsteinii
foliis rosulariis veris et supra minus hirsutis vel glabrescentibus, foliis caulinis minus numerosis, ab
Hieracio bifido
indumento plumoso differt.
Perennial, hemicryptophyte. Rhizome stout, vertical. Stem erect, vertical, cylindrical, stout (
2 mm
in diam.),
25–40 cm
tall, light green, brownish-purple at base, slightly striated, phyllopodous, in lower part with dense plumose whitish, soft, simple hairs and sparse minute (up to
0.1 mm
long) yellowish glandular hairs, above glabrous. Basal leaves 3–4, ovate to elliptical, blade 4–8 ×
3–4 cm
, greyish-green, at base rounded to shortly attenuate, entire or denticulate only in the lower part, petiole
1–3 cm
long, like the lower surface of leaves densely covered with
1–1.5 mm
long plumose whitish soft simple hairs, upper surface becoming glabrous or only covered with moderate to sparse plumose hairs, glandular and stellate hairs absent. Cauline leaves 2–3, lanceolate, entire, the lower one long attenuate in a short,
1 cm
long petiole, the one or two others attenuate-sessile, pubescence like on the basal leaves. Synflorescence racemose, peduncles 2–3,
3–7 cm
long, sparsely covered with stellate hairs, in the upper part with 1–2 filiform bracts
3–4 mm
long. Involucre almost ovoid,
8–9 mm
long. Involucral bracts in a few rows, dark green, linear-lanceolate,
1 mm
wide, acute, with dense stellate and few
0.3 mm
long glandular hairs, simple hairs lacking. Corolla limb ligulate, yellow, glabrous. Styles yellow. Margins of alveoli with short broad teeth. Achenes not seen.
Phenology:
—Flowering in June and July. Fruiting in July.
Distribution and ecology
:—
Hieracium moravense
is currently known from only the
type
locality in the massif of Mt. Moravë in the region of
Korcë
. This locality is found in the narrow rocky valley ending at the south-eastern border of the village of Drenovë. The few plants that we could observe were growing at
1030–1050 m
a.s.l. in fissures of steep, nearly vertical serpentine rocks on the hydrographic right of the gorge, mainly facing to the south-west (
Fig. 3
). Due to the difficult access to the site, only
two specimens
could be collected. This new hawkweed can be categorized as a metallophyte, and is also likely an obligate serpentinophyte, being able to tolerate the several chemical and physical anomalies of ultramafic soils, especially the elevated concentrations of trace metals such as Ni, Cr and Co.
Taxonomic affinities
:—As given in the diagnosis,
Hieracium moravense
is quite intermediate between
H. waldsteinii
subsp.
plumulosum
and
H. bifidum
. Such species with intermediate characters between a species of
Hieracium
sect.
Pannosa
and a species of
Hieracium
sect.
Hieracium
or
H.
sect.
Bifida
(Arvet-Touvet) A.R.Clapham
in
Clapham
et al.
(1952: 1135)
are well known from the Balkan Peninsula. They may have evolved by hybridisation. Examples are
Hieracium eriobasis
Freyn & Sintenis
in
Freyn (1897: 787)
(„pannosum – murorum“),
H. megalothecum
Zahn (1921: 587)
(„gymnocephalum – murorum“),
H. mattfeldianum
Zahn (1928: 383)
(„pannosum – bifidum“) and
H. wettsteinianum
Zahn (1921: 586)
(„gymnocephalum – bifidum“). In the habit they all are characterised in being intermediate between the aphyllopodous
Pannosa
species and the phyllopodous
H. murorum
Linnaeus (1753: 802)
or
H. bifidum
Kitaibel ex
Hornemann (1815: 761)
. Differences are easy to recognize by the different pubescence of the putative parents.
Final remarks
:—The discovery of
Hieracium moravense
underlies the phytogeographical importance of Mount Moravë as a centre of plant diversity in
Albania
, especially for the serpentine flora. This area is home of local endemics such as
Odontarrhena moravensis
(F.K.Meyer) L.
Cecchi & Selvi (2018: 18)
and
Centaurea drenovensis
Pils (2016: 9)
, as well as of Albanian endemics like
Acantholimon albanicum
O.Schwarz & F.K.Meyer
in
Meyer (1987: 31)
, and others.