Noideattella and Tolegnaro, Two New Genera of Goblin Spiders from Madagascar, with Comments on the Gamasomorphoid and Silhouettelloid Oonopids (Araneae, Oonopidae) Author Álvarez-Padilla, Fernando Author Ubick, Darrell Author Griswold, Charles E. text American Museum Novitates 2012 2012-05-25 2012 3745 1 76 http://www.bioone.org/doi/abs/10.1206/3745.2 journal article 2558 10.1206/3745.2 25e1eb0b-a2ad-4c0f-aaec-21d56c6fde5e 0003-0082 5360355 Noideattella assumptia ( Saaristo, 2001 ) , new combination Figures 31–60 , 372 Silhouettella assumptia Saaristo, 2001: 319 , figs. 32A, 33A, 34A, 35–39 ( paratypes examined from Seychelles , Aldabra , Assumption Atoll , 14 July 1984 , USSR Zool. Expedition, deposited in MZT, PBI_ OON 35298 ). DIAGNOSIS: N. assumptia can be differentiated from other Noideattella species by the following combination of characters: habitus orange-brown; carapace pars cephalica and sides strongly granulate (figs. 34, 41, 42 ), book lung covers large and very narrow (figs. 32, 33); sternum microsculpture surface finely reticulate distributed medially and in furrows (figs. 35, 43 ); female epigastric scutum A2 apodeme bases longer than 2 ⁄ 3 the genital opening length (figs. 39, 53). DESCRIPTION: Male (PBI_OON 3248): Total length 1.3. Cephalothorax: Carapace orangebrown. Sternum as long as wide, orange-brown, furrow with rows of small pits, anterior margin with interrupted transverse groove (figs. 35, 43, 45 ). Mouthparts: Chelicerae, endites, and labium orange-brown. Abdomen: ovoid. Dorsal scutum orange-brown, middle surface reticulate, sides reticulate. Postepigastric scutum orange-brown, long, semicircular, covering nearly full length of abdomen (figs. 31–33). Male genital opening slightly protruded, epigastric scutum apodemes small, contiguous with anterior book lung openings, connected by a furrow. Posterior spiracles connected by a groove (fig. 50). Legs: Pale orange, without color pattern. Leg spination (only surfaces bearing spines listed, all spines longer than segment width): tibiae: I p2-1-1; r2-1-0; II p2-2-1; r2-1-0; metatarsi: I, II p1-1-0; r1-1-0. Tarsal organ with four sensilla visible. Genitalia: Epigastric region with sperm pore situated at level of anterior spiracles (fig. 50). Palp proximal segments yellow; cymbium yellow, with distal patch of setae; bulb yellow (figs. 36–38). Embolus dorsal lamella with at least two distal apophyses (figs. 57–60), ventral lamella bifurcate (figs. 56, 59). Palp as in figures 55 and 56. Female (PBI_OON 3248): Total length 1.5. As in male except as noted. Abdomen: Postepigastric scutum not fused to epigastric scutum. Spinneret scutum with fringe of needlelike setae. Legs: Metatarsi III and IV weak ventral scopula present. Leg spination (only surfaces bearing spines listed, all spines longer than segment width): tibiae: I p2-1-1; r2-1-0; II p2-2-1; r2-1-0; metatarsi: I, II p1-1-0; r1-1-0. Trichobothria of metatarsus: I one, IV one. Tarsal organ number of sensilla variable, leg four has two sensilla, other tarsal organ sensilla vary from three to two. Genitalia: Genital opening slit shaped (fig. 51). Sperm sac secretory glands openings flat, concentrated around GAp process, GAp process shaped like twisted tube (fig. 54). Genitalia as in figures 39 and 40. VARIATION: Male paturon inner margin setae vary among specimens; some lack modified setae, while other specimens have a brush of modified seta (figs. 37, 47–49 ). Male leg II spination varies from three to four spines on the prolateral side of tibiae. Shape and number of embolar dorsal and ventral lamella apophyses shape varies among individuals. MATERIAL EXAMINED: N = 250. Madagascar : Antsiranana : Forêt Bekaraoka , 6.8 km 60° ENE Daraina , tropical dry forest, 150 m , S13.00277° , E49.01166° , Dec. 07, 2003 , Fisher-Griswold Arthropod Team , 5♂ ( CASC PBI _ OON 3958 ) ; Forêt d’Ampondrabe , 26.3 km 10° NNE Daraina , tropical dry forest, 175 m , S12.97000° , E49.70000° , Dec. 10, 2003 , Fisher-Griswold Arthropod team, 13 mixed ( CASC PBI _ OON 3579 ) ; Forêt d’Ampombofofo , littoral forest, 25 m , S12.09944° , E49.33861° , Nov. 21–22, 2007 , Fisher-Griswold Arthropod team, 1♂ ( CASC PBI _OON 36093) ; Forêt d’Orangea , 3.6 km 128° SE Ramena , littoral forest, 90 m , S12.25888° , E49.37472° , Feb. 22–28, 2001 , Fisher-Griswold Arthropod team, 30 mixed ( CASC PBI _ OON 3248 ) ; 1♂ ( CASC PBI _ OON 3250 ) ; 3♀ ( CASC PBI _ OON 3301 ) ; 1♀ ( CASC PBI _ OON 3932 ) ; Forêt di’Andavakoera, 21.4 km 356° N Betsiaka , rainforest, 425 m , S13.00194° , E49.00361° , Dec. 15, 2003 , Fisher-Griswold Arthropod team, 2 mixed ( CASC PBI _ OON 3956 ) ; Montagne des Français , 7.2 km 142° SE Antsiranana (= Diego Suarez ), tropical dry forest, 180 m , S12.32277° , E49.33805° , Feb. 22–28, 2001 , Fisher-Griswold Arthropod team, 6 mixed ( CASC PBI _ OON 3264 ) ; 5 mixed ( CASC PBI _ OON 3273 ) ; 1♀ ( CASC PBI _ OON 3930 ) ; Nosy Be , Lokobe Forest , S13.41633° , E48.30736° , Aug. 11–14, 1992 , V. Roth , B. Roth , 1♂ ( CASC PBI _ OON 3949 ) ; Parc National Montagne d’Ambre , 3.6 km 235° SW Joffreville , tropical dry forest, 925 m , S12.53444° , E49.17944° , Jan. 20–26, 2001 , Fisher-Griswold Arthropod Team , 1♂ ( CASC PBI _ OON 3268 ) ; Reserve Analamerana , 16.7 km 123° SW Anivorano Nord , tropical dry forest, 225 m , S12.67138° , E49.37388° , Dec. 03, 2004 , Fisher-Griswold Arthropod team, 9 mixed ( CASC PBI _OON 35275) ; 5 mixed ( CASC PBI _OON 35264) ; Reserve Speciale d’Ambre , 3.5 km 235° SW Sakaramy , 325 m , S12.46888° , E49.24222° , Jan. 26–31, 2001 , L.J. Boutin , 2 mixed ( CASC PBI _ OON 3261 ) ; Reserve Speciale d’Ambre , 3.5 km 235° SW Sakaramy , tropical dry forest, 325 m , S12.46888° , E49.24222° , Jan. 26–31, 2001 , Fisher-Griswold Arthropod Team , 18 mixed ( CASC PBI _ OON 3263 ) ; 18 mixed ( CASC PBI _ OON 3276 ) ; 4♀ ( CASC PBI _ OON 3963 ) ; 2 mixed ( CASC PBI _OON 36270) ; Reserve Speciale de l’Ankarana , 22.9 km 224° SW Anivorano Nord , tropical dry forest, 80 m , S12.90888° , E49.10972° , Feb. 10–16, 2001 , Fisher-Griswold Arthropod Team , 1♀ ( CASC PBI _ OON 3285 ) ; 1♀ ( CASC PBI _ OON 3933 ) ; 16 mixed ( CASC PBI _ OON 3957 ) ; 4♀ ( CASC PBI _ OON 3960 ) ; 5 mixed ( CASC PBI _OON 36234) ; Reserve Speciale l’Ankarana , 13.6 km 192° SSW Anivorano Nord , tropical dry forest, 210 m , S12.86361° , E49.22583° , Feb. 16–20, 2001 , Fisher-Griswold Arthropod Team , 1♂ ( CASC PBI _ OON 3241 ) ; 24 mixed ( CASC PBI _ OON 3242 ) ; Reserve Speciale l’Ankarana , 22.9 km 224° SW Anivorano Nord , Camp Anglaise , 80 m , S12.90888° , E49.10972° , Feb. 10–16, 2001 , L.J. Boutin , 1♀ ( CASC PBI _ OON 3297 ) . Seychelles : Aldabra , Assumption Atoll , July 14, 1984 , USSR Zool. Expedition , 2 mixed, paratypes ( MZT PBI _OON 35298) .