Three new species of Tracholena Common, 1965 (Lepidoptera: Tortricidae: Tortricinae: Schoenotenini) from New Caledonia associated with Araucariaceae Author Dugdale, John S. text Zootaxa 2005 870 1 16 journal article 10.5281/zenodo.170880 4302257a-8b19-4c5e-ac2e-0256b9294ca1 1175­5326 170880 Tracholena liparodes sp. nov. ( Figs. 2 , 6, 9, 10 , 13 ) Material examined : Holotype . Male, “NOUVELLE CALEDONIE: Mt Panié, 1380 m to light, 15 December 1990 , J.S. Dugdale.” “ HOLOTYPE male/ Tracholena liparodes Dugdale ” [red card], MNHN Paratypes . New Caledonia : 1 male , 1 female , same data as holotype except capture dates span 6–16 December 1990 , ONNC Noumea; 1 male , 1 female , ditto, NZAC . Description : Male, female ( Fig. 2 ): Head with vertex tufts grey brown, scape grey brown; with vom Raths organ less than half length of labial palp apical segment; maxillary palp one­segmented on an inwardly spinulose pedicel, apical palp segment with 1–2 scales and apically with 4 short stiff sensilla. Flagellomeres longer than wide, 1–3 cylindrical, rest tapering apically, and each with a basal whorl of trichomes longest on the basal flagellomeres and at most only slightly longer than flagellomere length, flagellomeres with the paired sensilla coeloconica contiguous, ventro­apical; labial palpus second segment scaling evenly grey­brown speckled outwardly; prosternal midline scale tuft pale buff contrasting with the brown speckled fore coxae. Forewing: Shape more or less as in paniense ( Fig. 2 ), length ca. 2.3X width (females), ca. 2.8X width (males); forewing underside with costal cell area dark grey­scaled, contrasting with the rest of the wing; forewing upperside (and mesonotum) ochreous grey, with indistinct blackish markings at about ¼, ½, and a series marking apices of veins R1, R2, and R3; an interrupted serrate dark brown­scaled line at ¼, and another, even less complete at about ½ wing length; Sc, R and branches with scattered small groups of dark brown scales, 1A+2A; discally with scattered irregular patches of more ochreous scales; areas of raised scales as in nigrilinea . Hindwing: Grey buff; the anal tuft over half the length of anal wing margin; venation: veins M3 and Cu1A coincident at base (i.e., “bifid”) and distant from vein M2 ( Fig. 6 , cf. Fig. 5 ). Wingspan : Holotype male: 15.2 mm ; 2 paratype males: 14.8 and 15.2 mm ; paratype females 15.8, 16.0 and 18.2 mm , respectively. Male genitalia ( Figs. 9, 10 ): Uncus long, slender, decurved; uncus base broad, mesal part as in nigrilinea , produced anteriorly between the posteriorly extended tegumen apices; socii lobate; gnathos arms forming a U thickened towards apex, with lateral spinules outwardly, and thickened part with dorsal spinules, apex with a pair of ventral “wings; transtilla dorsally with a pair of dorsally and outwardly­directed spinose clavate arms; valva apex oblique (costal margin shorter than saccular margin), and no sclerotised strut below the M2 muscle attachment ( cf. nigrilinea ); aedeagus ( Fig. 9 ) stout, not longer than valva; with a large apical dextral hook (appressed to the vesica in Fig. 9 ); bulbus ejaculatorius entering at ca. half aedeagus length, vesica with a basal spinulose patch and 14 nondeciduous plate­like cornuti arranged in 2 arcs; bulbus ejaculatorius tube stout, sinuous, hood not as long as sleeve. Female genitalia ( Fig. 13 ): Ovipositor lobes axes more or less parallel, unmodified anteriorly. Ovipore on a level with anterior third of lobes, and minutely spinulose; sternite 8 with a raised pair of transverse curved ridges; lamella postvaginalis a broad scobinate field with a narrow, sclerotised, scobinate strip bordering the ostium bursae; lamella antevaginalis a thin sinuous sclerotised bar, ostium bursae laterally produced anteriorly (but no ligament discernible); colliculum short, ventrally split; ductus seminalis arising on a latero­ventral papilla; ductus bursae and cestum sinuous, forming one long complete spiral; corpus bursae with a broad­margined, thorn­like signum, corpus bursae covered in minute pit­like micro­sculpture. Diagnosis: This species resembles T. paniense and T. lipara , being broad­winged (forewing length <2.9X width). It can be distinguished from those two by the forewing with costal cell area dark­grey scaled ventrally (not so in the others); hindwing veins M3 and CuA1 contiguous basally but not stalked (stalked in lipara , distant in paniense ); aedeagus with an apical hooked process; and vesica with 14 chunky, non­deciduous, basal cornuti (a possible species apomorphy). The unmodified ovipositor lobes, the sinuous cestum/ductus bursae (a single, elongate spiral), and presence of a strong hollow signum are character states shared with the Queensland T. micropolia . Other differences are noted in the “Diagnosis” of T. paniense below. General notes : While most specimens examined have a wingspan of 14–16 mm , one female of this species exceeds 18 mm , but it has the same venation and genital conformation as the smaller specimens of liparodes . Similar variation in wingspan was reported for the female­strobilus/seed­eating T. lipara ( Common 1973: 302 ) but not for the foliage­eating T. dialeuca and T. hedraea ( Common 1982: 222, 223 ), nor for the male­strobilus­eating T. nigrilinea (see above). Specimens were collected at light at the same time and at the same site as T. paniense , under a stand of Agathis montana . No feeding site on any host could be found. Most specimens are greasy, indicating that larvae may be seed­eaters. Common (1973: 303) noted that T. lipara specimens, reared from damaged klinkii pine ( Araucaria hunsteinii ) seeds, can become greasy, and most T. nigrilinea specimens, all from male cones, are greasy. Superficially, this species resembles 2 unset T. lipara specimens (in ANIC, CSIRO) from Bulolo in the Papua New Guinean Highlands, but lacks the transverse lines of orange scales so characteristic of that species. Etymology: The name refers to the external resemblance to T. lipara .