New species and records of Copris (Coleoptera: Scarabaeidae; Scarabaeinae) from Central America Author Kohlmann, Bert Author Delgado, Enio Cano And Leonardo text Zootaxa 2003 167 1 16 journal article 10.5281/zenodo.156784 7450e6f5-2d35-43f7-9cbc-5aa4eab735b6 1175­5326 156784 Key to the species of the remotus complex based on the major males 1. Head unarmed having only a gibbosity, clypeus tridentate; thorax unarmed. Costa Rica ........................................................................................ tridentatus Solís & Kohlmann ­ Head armed with a horn, clypeus bidentate; thorax armed .......................................... 2 2. Pygidial margin incomplete, its inner border completely effaced ventrally. Chiapas, Guatemala , Costa Rica and Panamá .............................................. costaricencis Gahan ­ Pygidial margin complete, its inner border entirely engraved ...................................... 3 3. Eighth elytral stria incomplete; frontal pronotal declivity with a carina running through its middle ....................................................................................................... 4 ­ Eighth elytral stria complete; frontal pronotal declivity without a carina running through its middle ....................................................................................................... 5 4. Antennae dark­brown; male anterolateral pronotal angles not forming median point, but presenting an excavation behind these angles ( Fig. 4 ), median pronotal prominences diverging ( Fig. 1 ). Guatemala . caliginosus Kohlmann, Cano & Delgado , n. sp. ­ Antennae reddish­brown; male anterolateral pronotal angles forming a small downwardly directed median point and separated from the angles by a more or less level area ( Fig. 4 ), median pronotal prominences strongly diverging ( Fig. 2 ). Guatemala and Honduras ............................................. nubilosus Kohlmann, Cano and Delgado, n. sp. 5. Protibial apical spur abruptly bent inward very near apex. Oklahoma, Texas, Coahuila, Nuevo León and Tamaulipas .............................................................. remotus LeConte ­ Protibial apical spur curved or slightly curved inward ( Fig. 3 ) ................................... 6 6. Anterior pronotal margin with a minute, acute median tooth ( Fig. 3 ); elytral striae with large, less transverse punctures, distinctly crenating the margins of the strial channels; protibial apical spurs blunt ( Fig. 3 ). Veracruz to Chiapas ........................ sallei Harold ­ Anterior pronotal margin not forming any median point or angle; elytral striae with fine, very transverse punctures which at most only slightly crenate the edges of the channel; protibial apical spurs pointed. Michoacán .. mexicanus Matthews and Halffter