A new record of Neoperla obliqua Banks, 1930 (Plecoptera: Perlidae) from Mt. Malindang, Mindanao, Philippines and association of life stages using DNA barcodes
Author
Dela Cruz, Ian Niel B.
Author
Nuñeza, Olga M.
Author
Lin, Chung-Ping
text
Zootaxa
2018
2018-11-06
4514
1
145
150
journal article
28106
10.11646/zootaxa.4514.1.12
235da605-d088-41da-98ce-5e9c7d141a47
1175-5326
2605782
F2CAE9F9-2975-48AF-868D-11173F16ED48
Neoperla obliqua
,
Banks 1913
Material examined.
Philippines
,
Mindanao
,
Mt. Malindang
,
Layawan River
.
1 ♂
,
127masl
,
N 08°26.314’
E 123°42.492’
,
5 October 2013
;
3 ♀
,
185masl
,
N 08°25.418’
E 123°41.806’
,
3 October
2013
. 185masl,
N 08°25.418’
E 123°41.806’
,
3 October 2013
;
1 ♀
nymph,
1,218masl
,
N 08°18.495’
E 123°37.980’
. All material is deposited in the Natural Science Museum, MSU-IIT,
Iligan City
,
Philippines
.
Ova.
Shape oval, highly striated, (
Fig. 1A
). Collar present but flat. Length ca. 318.16 ± 10.37 µm (n=3), equatorial width ca. 222.63 ± 4.97 µm. Collar bare, with length ca. 16.2 ± 8.5 µm and width ca. 85.83 ± 9.7 µm, with uneven surface, resembling a lid (
Fig. 1C
). Chorionic surface rugose and somewhat ornate. Gyral structure irregular but usually follows a longitudinal ridge-like pattern which arises from the collar end, creating a sulci or irregular grooves in between studded with pores of almost similar sizes (
Fig. 1D
). Micropyles at distal end of the collar and present in all ova examined (
Fig. 1B
).
Nymph.
General color golden ochre to pale orange, while darker, usually dark brown to black margins on frons and labrum of head, the pronotal disc, tip of wingpads, and abdominal terga (
Fig. 2A
). Pronotal disc ovalshaped. Wingpads not elongated. Terga faintly striped. Antennae and cerci setaceous. Mandible ochre, ovateshaped but broad on its base possessing a five comb-like, blunted teeth of irregular sizes (
Fig. 2B
). Few fine setal hairs on the inner edges of mandible. Legs golden-brown without acute setal surface hairs (
Fig. 2C
).
FIGURE 1.
Neoperla obliqua
ova. (1A) Entire ova. (1B) Chorionic details (arrow refers to micropyle). (1C) Collar end. (1D) Anterior end. Scale = 100µm.
Female nymph.
Body length of
17.44 mm
(n=1), abdominal length
7.02mm
. Width of head is
4.01mm
, pronotal disc is
3.91mm
. Ocelli
0.8mm
diameters apart. Mandible
1.43 mm
. Mesothoracic and metathoracic width
4.36 mm
and
4.07 mm
, respectively. Antenna
11.05 mm
. Femur
4.13 mm
, tibia
3.94 mm
, tarsus
1.22 mm
. Cercus
9.78 mm
.
Habitat
. Adults of
N. obliqua
were sympatric with another unidentified perlid stonefly species. The single nymph of
N. obliqua
was found under large rocks in the Layawan River, in a section with moderate current.
Life stage and sex association.
Pairwise
CO1
sequence comparisons revealed 0.2% intraspecific divergence between males and females of putative conspecifics. The sequence similarity between adult and nymphal stages was 99.9 ± 0.14% (
Table 2
). Comparisons between divergences of interspecific
CO1
sequences of all available perlid stoneflies based on Kimura-2-parameter (K2P) model were also shown (
Table 3
).