Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula
Author
Mertlik, Josef
Pohřebačka 34, CZ- 53345 Opatovice nad Labem, Czech Republic
mertlik@elateridae.com
Author
Németh, Tamás
Hungarian Natural History Museum, Department of Zoology, Baross u. 13, H- 1088 Budapest, Hungary
haesito@gmail.com
Author
Kundrata, Robin
Department of Zoology, Faculty of Science, Palacky University, 17. listopadu 50, CZ- 77146 Olomouc, Czech Republic
robin.kundrata@upol.cz
text
Zootaxa
2017
2017-01-13
4220
1
1
63
journal article
7453
10.11646/zootaxa.4220.1.1
eb597d59-1f6e-412d-a99a-de1a67e7cc94
1175-5326
4670787
D74BC90C-84CC-4788-9048-54F5C8521B32
Dima assingi
Schimmel & Platia, 2008
Figs 6–9
,
120
,
175–177
,
246
.
Dima assingi
Schimmel & Platia, 2008
: 572
.
Dima peloponnensis
Schimmel & Platia, 2008
: 586
,
syn. n.
Type depositories.
Holotype of
D. assingi
, ♂ (PCRS); paratype of
D. assingi
, ♀ (PCRS); holotype of
D. peloponnensis
, ♂ (PCRS); 2 paratypes of
D. peloponnensis
, ♀♀ (PCRS, PCGP).
Type
locality.
Greece
:
Peloponnese
,
Aroania Mts.
,
1450 m
,
10 km
E of Kalavrita
(for both
D
.
assingi
and
D
.
peloponnensis
).
Redetermined
material.
GREECE
:
Morea
,
Trikala
,
Kilini Mts.
, 1.–9.
VI.1996, 1
♂,
1 ♀
,
Padovani
&
Malmusi
leg. (as
D. hladilorum
in
Schimmel & Platia 2008
) (
PCGP
)
;
Peloponnese
,
Kilini Mts.
,
Trikala
,
1100 m
, 29.V.–7.
VI.1998, 1
♀,
L. Saltini
leg. (as
D. isabellae
Dajoz,
1973
in Schimmel & Platia
2 008) (
PCRS
)
;
Peloponnese
,
Korinthos
:
Stilia
,
700 m
(=
Kilini Mts.
), 30.V.–6.
VI.1998, 1
♂, leg.
L. Saltini
leg. (as
D. raineri
Wurst,
1997
in
Schimmel & Platia 2008
) (
PCRS
)
.
New
material.
GREECE
: North
Peloponnese
[Nordpeloponnes],
Xylokastro
,
Zemeno
env.,
Driodasos Mongostou
(ca.
37°59'47.42"N
,
22°35'43.23"E
), 28.V.–7.
VI.1997, 1
♀,
Manfred Egger
leg. (
PCME
).
Diagnosis.
Dima assingi
is a large, moderately convex species (
13.5–16.3 mm
long), with pronotum with coarse punctation, long straight semi-erect to erect pubescence on the sides (
Fig. 120
), and elytra with decumbent to semi-erect pubescence basally.
Dima assingi
is very like
D
.
neumanni
Platia, 2013
(
Fig. 67–74
) and
D
.
schnitteri
Platia, 2013
(
Fig. 105–108
), which also occur in the northern
Peloponnese
. They are characteristic by the long straight semi-erect to erect setae on the pronotal sides and semi-erect pubescence on elytra. Together they form a complex of morphologically variable species with unclear specific limits.
Dima assingi
differs from two above mentioned species in elytra with shorter decumbent to semi-erect pubescence in basal third or half, and in semierect setae in the first half of pronotal sides (
D
.
neumanni
and
D
.
schnitteri
have longer semi-erect to erect setae in basal third or half of elytra and erect setae in the anterior half of pronotal lateral margin;
Figs 120
,
146
,
157
). In contrast to the typical
D. neumanni
and
D. schnitteri
, all known specimens of
D. assingi
have relatively long pronotum, however, some specimens of
D. neumanni
(mainly from Aroania Mts.) have similarly shaped pronota.
Intraspecific variability.
There is a variability in the coloration of body and pubescence (the specimens from Aroania Mts. have paler pubescence), pronotal pubescence orientation, and in the shape of apical lobe of paramera (
Figs 6–9
,
175–177
).
Distribution.
Greece
(
Peloponnese
: Aroania Mts., Kilini Mts.;
Fig. 246
).
Remarks.
The
type
specimens of
D
.
assingi
and
D
.
peloponnensis
were collected at the same time and on the same locality in Aroania Mts. (
Schimmel & Platia 2008
). They differ only in the body coloration (
D
.
peloponnensis
is slightly darker;
Figs 6–7
), surface of pronotum (wrinkled, with different orientation of setae in
D
.
peloponnensis
), and the shape of an apical lobe of paramera (shorter in
D
.
peloponnensis
;
Fig. 177
). All these characters are variable in
Dima
species. The specimens of
D
.
assingi
from the Kilini Mts. are similarly colored as
D
.
peloponnensis
and some of them vary slightly in the orientation of setae in the pronotum. The wrinkled pronotum is probably a molting defect known also in other species (see e.g.
D
.
fialai
sp. nov.
or
D
.
fthiotidensis
Schimmel & Platia, 2008
). Therefore, we herein synonymize
Dima peloponnensis
Schimmel & Platia, 2008
under
Dima assingi
Schimmel & Platia, 2008
.
There are two species of
Dima
reported from the Aroania Mts., i.e.
D
.
assingi
and
D. neumanni
(
Fig. 246
). Both probably differ in their habitat preference. Specimens of
D. assingi
from the Kilini Mts (Zemeno, Stilia, Trikala) were found near seasonal streams in agricultural lands, whilst
Dima neumanni
was collected in the dark wet gorges in the fir forests of the Aroania and Erimanthos Mts. It must be also noted, that the occurrence of
D
.
assingi
in Aroania Mts.
was not confirmed by the recent field research by JM and colleagues—only
D
.
neumanni
were found in the vicinity of the
type
locality of
D
.
assingi
. We cannot exclude a possibility that the
type
specimens of
D. assingi
were mislabelled.