Revision of Taurocerus (Heteroptera, Pentatomidae, Pentatomini)
Author
Jocélia Grazia
Departamento de Zoologia, Universidade Federal do Rio Grande do Sul, Av. Bento Gonçalves, 9500, Bloco IV, Prédio 43435, 91501 - 970 Porto Alegre, RS, Brazil.
jocelia@ufrgs.br
Author
Aline Barcellos
Museu de Ciências Naturais, Fundação Zoobotânica do Rio Grande do Sul, Caixa Postal 1188, 90001 - 970 Porto Alegre, RS, Brazil.
alinebar@fzb.rs.gov.br
text
Iheringia, Série Zoologia
2005
2005-06-30
95
2
173
181
journal article
10.1590/S0073-47212005000200007
a20dc7bd-2f44-4d2e-a5e6-bfd99c1f3b77
3630712
Taurocerus edessoides
(
Spinola, 1837
)
(
Figs. 6, 9, 12, 15, 18, 21, 24, 26, 28
)
Arvelius edessoides
SPINOLA, 1837:347
.
Taurocerus hector
STÅL, 1862:108
;
1872:47
;
WALKER, 1867:391
(list).
Taurocerus edessoides
;
DALLAS, 1851:302
(list);
DISTANT, 1880:82
, pl. 7, fig. 25 (list);
Stål, 1872:47
;
LETHIERRY & SEVERIN, 1893:185
(cat.);
KIRKALDY, 1909:150
(cat.).
Type-locality. Brazil.
Type
material.
Lectotype
and
paralectotype
, here designated. The material belongs to Spinola’s collection, at present deposited at MRSN. A green label in the box, containing the following data:
Arvelius edessoides
Spin
., G.
taurocerus
, A. et S., D. Bouquet, Brésil.
Type
material of
T. hector
examined. Stål described this species based on two males from Mexico and one female from unknown locality. We received from NHRS three males labeled as “
typus
” (loan number 1068/99) and “
paratypus
” (numbers 1069/99 and 1070/99), the last one collected in Bogotá [Colombia] by Alexander Lindig. We believe that the specimen 1070/99 is not actually a
syntype
of
T. hector
and the female mentioned in the Stål’s original description is missed. Then, we here designate as
lectotype
and
paralectotype
, respectively: 1, labeled: a) Vera Cruz [handwritten]; b)
Hector
Typ. Stål [handwritten]; c) Sallé; d)
Typus
[red label]; e) Naturhistoriska Riksmuseet Stockholm Loan no 1068/99; 1, labeled: a) Mexico; b) Stål; c)
Hector
Typ. Stål [handwriting]; d)
Paratypus
; e) Naturhistoriska Riksmuseet Stockholm Loan number 1069/99.
Description. Dorsal surface reddish-castaneous, abdominal venter reddish. Apex of humeri yellow. Rostrum reaching the fourth abdomninal sternite. Vshaped callous of scutellum, in the most of the specimens examined, with broad and closed arms, becoming cordiform in appearance; in some specimens, however, the callous is faded.
Male. Measurements (n = 10). Total length 15.56 (14.63-16.50) ± 0.73; head length 2.96 (2.69-3.16) ± 0.13; width 3.27 (3.06-3.50) ± 0.15; eye width 0.74 (0.63-0.81) ± 0.06; interocular distance 1.80 (1.69-1.94) ± 0.08; anteocular length 1.47 (1.38-1.56) ± 0.07; antennal segments I, 0.88 (0.81-1.06) ± 0.08; II, 1.47 (1.25-1.63) ± 0.10; III, 1.94 (1.63-2.44) ± 0.22; IV, 3.13 (2.78-3.56) ± 0.22; V, 3.24 (3.06-3.44) ± 0.14; pronotal length 3.81 (3.63- 4.25) ± 0.24; width 13.62 (11.88-15.00) ± 1.19; scutellum length 6.70 (6.25-7.25) ± 0.36; width 5.98 (5.50-6.50) ± 0.35; corium length 9.34 (8.50-10.00) ± 0.50; abdominal width 9.24 (8.75-9.63) ± 0.35.
Genitalia. Pygophore quadrangular, globose in caudal view, little open dorsally. Dorsal rim slightly sinuated. Proctiger trapezoidal, wider basally, with 1 + 1 tumid areas at the distal half (
fig. 6
). Parameres distally thin (
fig. 15
), apex strongly curved latero-posteriorly (
fig. 9
). Posterolateral angles with a small, acute tubercle. Ventral rim: median process acute; lateromedian processes small, conical, delimiting a median V-shaped excavation (
fig. 12
).
Phallus
(
figs. 18, 21, 24
): basal plate welldeveloped, half as long as
phallotheca.
Visible part of
processus phallothecae
long, more than two thirds as long as
phallotheca
. Vesica well-developed with one median, ventral process.
Ductus seminis distalis
long, surpassing conjunctiva. Conjunctiva well-developed, with 1 + 1 dorsal processes (prcj1); prcj2 absent.
Female. Measurements (n = 10). Total length 15.56 (14.25-17.00) ± 0.98; head length 2.87 (2.66-3.19) ± 0.13; width 3.28 (3.13-3.44) ± 0.10; eye width 0.77 (0.69-0.81) ± 0.04; interocular distance 1.80 (1.69-1.88) ± 0.08; anteocular length 1.43 (1.31-1.56) ± 0.07; antennal segments I, 0.92 (0.81-1.06) ± 0.07; II, 1.40 (1.25-1.56) ± 0.10; III, 1.83 (1.63-2.00) ± 0.11; IV, 3.06 (2.81-3.25) ± 0.15; V, 3.18 (3.06-3.25) ± 0.10; pronotal length 3.81 (3.56-4.00) ± 0.17; width 13.41 (12.13-14.75) ± 0.78; scutellum length 6.73 (6.25-7.19) ± 0.30; width 6.01 (5.75-6.38) ± 0.21; corium length 9.31 (8.88-10.00) ± 0.42; abdominal width 9.29 (8.88-9.75) ± 0.32.
Genitalia. Gonocoxites 8 triangular, about one and a half wider than long (
fig. 26
). Sutural borders of gonocoxites 8 convergent, contiguous at apical half. Posterior borders concave near laterotergites 9. Laterotergites 9 narrow at apex. Secondary thickenings of gonapophysis 9 ear-like (
fig. 28
).
Material examined.
MEXICO
,
Veracruz
:
playa escondida 28 Km NE of Catemaco
,
50 m
),,
30.IX.
[19]74
,
M. Madison
col. (
AMNH
);
Los Tuxtlas
(“
camino a Balzapotl
”),,
11.IX.
[19]86
,
L. Cervantes
col.(10:45 80,
Siparuna
) (
AMNH
);
idem,,
15.VI.
[19]86
, idem (13:20,
Siparuna
), idem (
AMNH
);
GUATEMALA
,
Alta Verapaz
:
San Juan,
,
Champion
col. (
NHMW
)
;
COSTA RICA
,
Guanacaste
:
Santa Rosa
(PN Guanacaste
),,
20.X.
[19]96
,
F. Fernández
col. (IAvH 05098)
;
PANAMA
,
Panama
:
Distr. Chame
,
Cerro Campana
(
800 m
),,
24.VII.
[19]82
,
H.D. Engleman
col. (
AMNH
)
;
idem,,
3.V.
[19]81
, idem (
AMNH
)
;
idem,,
10.V.
[19]81
,
E. Giesbert
col. (
AMNH
)
;
Canal Zone
:
Barro Colorado
,,
II-III.1936
,
W.J. Gertsch
col. (
AMNH
)
;
COLÔMBIA
,
Meta
:
Caño Grande
(
450 m
),,
II.[19]48
,
Richter
col. (
UNCB
He 00435)
;
Amazonas
:
Araracuara
,,,
21.VIII.
[19]77
,
R. Restrepo
col. (
UNCB
He 00376
,
He 00377
)
;
ECUADOR
,
Napo
:
Limoncocha
(
00 o 24’S
,
76 o 36’W
),,
7.VIII.1972
,
Peter L. Kazan
col. (
AMNH
)
;
GUYANA
,
Cuyuni- Mazaruni
:
Bartica
,, VIII,
Pomerat
col. (
AMNH
)
;
BRAZIL
, C. Samuel,
Jamari
,,
III.1944
,
Parko
col. (
UFRG
)
;
Amazonas
:
Benjamin Constant
(
Rio Javari
),,
III.1942
,
August Rabaut
col. (
AMNH
)
;
idem,,
25.I-15.II.1942
(
AMNH
)
;
Querari
(
Pelotão
),
01 o 05’N
/
69 o 51’W
,,
20-21.IV.1993
,
C. Motta
,
R. Ferreira
,
J. Vidal
,
B. Matteo
col. (“
luz mista de mercúrio e luz negra
BL
e
BLB
”) (
INPA
)
;
Pará
:
Belém
(
Utinga
),,
20.I.1977
,
N. Guimarães
col. (
MPEG
)
;
Acre
:
Feijó
,, 2,
XI.1956
(
MZSP
)
;
idem,,
XII.1956
(
MZSP
)
;
Mato Grosso
:
Chapada [dos Guimarães]
,, IX, acc.13739 (
AMNH
)
;
idem,, VIII, acc. 23739 (
AMNH
)
;
Rondônia
:
Porto Velho
(
Terr. Guaporé
),,
X.[19]44
,
Parko
col. (
UFRG
)
;
Vilhena
,,
11.XII.1986
,
C. Elias
col.,
Polonoroeste
(
DZUP
)
;
Rolim de Moura
(
Linha 204
),,
31.X.1978
,
K. S. Brown Jr.
col. (“
em mato aberto, terra roxa
”) (
ZUEC
2544
)
;
Minas Gerais
:
C[armo do] R[io] Claro
,,
I.[19]45
,
Carvalho
col. (
MNRJ
)
;
Espírito Santo
:
Baixo Guandu
,,
23-30.XI.1970
,
C. Elias
col. (
DZUP
)
;
Santa Leopoldina
,,
3.IX.1964
,
Claudionor Elias
col. (
DZUP
)
;
São Paulo
:
Tupã
,,
V.1947
,
Fritz Plaumann
col., Ex. Coll. H. Ruckes (
AMNH
)
;
Severinia
,,
XII.1940
,
A.G. Silva
col. (
MCNZ
003616
)
;
Osasco
,,
XII.1957
(
MZSP
)
;
PERU
,
Loreto
:
Iquitos
,, no date, F6062, H. Bassler Collection, Acc. 33591 (
AMNH
)
;
Upper Rio Maranon
,,
20.IV.
[19]24
, F6029, idem (
AMNH
)
;
idem,,
5.X.
[19]24
, F6103, H. Bassler Collection, Acc. 33591 (
AMNH
)
;
idem, 3,
11.X.
[19]24
, F6102, idem (
AMNH
)
;
idem,,
11.X.
[19]24
, F6102, idem (
AMNH
)
;
Amazonas
:
Rio Santiago
,,
06.XI.1928
, F619F, H. Bassler Collection, Acc. 33591 (
AMNH
)
;
idem,,
27.XI.
[19]24
, F6012, idem (
AMNH
)
;
idem,,
23.XI.
[19]24
, F6012, idem (
AMNH
)
;
Rio Ucayali
,,,
27.IX.
[19]23
, F6004, idem (
AMNH
)
;
Ucayali
:
Pucallpa
,
Rio Ucayali
,
200m
,, no date,
Dirings
col., Coll. Dirings (
MCNZ
003585
)
;
Huánuco
:
Tingo María
,,
18-23.IX.
[19]76
,
Jeff Baier
col. (
AMNH
)
;
Cuzco
:
Pilcopata
,,
11.X.68
, (“arbusto, 1604”) (
UFRG
)
;
BOLIVIA
,
Cochabamba
:
Chaparé
(
400 m
),,
12.XII.1949
,
Dirings
col. (
MZSP
)
.
Figs. 25-26. Genital plates, ventral view: 25,
T. achilles
Stål, 1862
; 26,
T. edessoides
(
Spinola, 1837
)
(gc8, gonocoxite 8; gc9; gonocoxites 9; la8, laterotergite 8; la9, laterotergite 9; VII, seventh abdominal sternite; X, tenth segment). Scale bar, 1 mm.
Figs. 27-28. Laterotergites, gonocoxites and gonapophyses of ninth segment, and ectodermical genital ducts, ventral view: 27,
T. achilles
Stål, 1862
; 28,
T. edessoides
(
Spinola, 1837
)
(aac, anterior annular crest; dr,
ductus receptaculi
; cs,
capsula
seminalis; gc9, gonocoxites 9; la9, laterotergite 9; mt, median thickening of gonapophyses 9; pac, posterior annular crest; pi,
pars intermedialis
; st, secondary thickenings of gonapophyses 9; tvi, thickening of vaginal intima; X, tenth segment). Scale bar, 0.5 mm.
Comments. The color of the specimens varies from reddish to dark reddish-castaneous; the length of humeri is also variable, but the apices are always yellowish. The size and the triangular shape of gonocoxites 8, combined with the color of humeri distinguish
T. edessoides
from
T. achilles
and
T. abruptus
.