New archaeorthopteran insects from the Late Carboniferous of the Nord and Pasde-Calais basins in northern France (Insecta: Cnemidolestodea, Panorthoptera)
Author
Coty, David
Author
Háva, Jiří
Author
Prokop, Jakub
Author
Roques, Patrick
Author
Nel, André
text
Zootaxa
2014
3878
5
462
470
journal article
42309
10.11646/zootaxa.3878.5.4
697c91ee-9d8b-4095-bd15-e55709b2b6bd
1175-5326
229037
377F9037-D039-44A5-B562-051560CBC9E0
Cf.
Tococladus
species undetermined
(
Fig. 1
)
Material studied:
Specimen 0 0 13 (print and counterprint), coll. Patrick Roques, to be deposited in the Musée Géologique Pierre Vetter, Decazeville,
France
.
Age and outcrop
.
Type
locality and horizon. Moscovian (Westphalian C/D equivalent to Bolsovian/Asturian), ‘Terril N°7’, Avion, Pas-de-Calais,
France
.
Description
. A fragment of wing corresponding to the anterior part of the apex, fragment
19.4 mm
long,
11.7 mm
wide; convex RA with a series of sigmoidal anterior veinlets, RA distally fused with concave main branch of
RP
, near wing apex, common branch
2.6 mm
long; a series of straight crossveins in area between RA and
RP
;
RP
with seven to nine parallel posterior branches with a series of simple straight crossveins between them; apical parts of branches of M and possibly CuA also preserved, parallel to the branches of
RP
.
Discussion
. Although very incomplete, this wing fragment fits quite well with the equivalent portion of a forewing of
Tococladus ralus
Carpenter, 1966
,
type
species of the Permian genus
Tococladus
. It is especially remarkable in the distal fusion of RA with
RP
near wing apex and the numerous parallel posterior branches of
RP
with short crossveins between them (see Béthoux et al., 2003: fig. 1). Of course an attribution to the genus
Tococladus
sensu stricto
is impossible to establish. The main interest of this fossil is that it represents the second
Cnemidolestodea
from the outcrop of Avion, the first one being
Aviocladus pectinatus
Prokop, Roques and Nel, 2014
, from which it differs in the more numerous branches of
RP
and veinlets between RA and costal margin more sigmoidal (
Prokop et al., 2014
).