Phylogenetic Classification of the Atlides Section of the Eumaeini (Lepidoptera, Lycaenidae) Author Martins, Ananda Regina P. Author Duarte, Marcelo Author Robbins, Robert K. text Zootaxa 2019 2019-02-28 4563 1 119 134 journal article 28153 10.11646/zootaxa.4563.1.6 0d849a4d-5dd1-4f60-9036-939f4486fe72 1175-5326 2601017 8E2935BD-3608-4BE1-B565-AE50966EE510 Brangas Hübner , [1819] type species: Papilio caranus Stoll Dabreras Bálint, 2008 , New Synonym type species: Thecla teucria Hewitson Brangas is characterized by eight synapomorphies ( Table 1 ), but they are exceedingly tentative because we included only two Brangas species in the phylogenetic analyses. However, every Brangas species has a male genitalia dorsal vinculum process that lies under and inside of its associated brush organs, a trait that occurs nowhere else in the Eumaeini . There are sixteen described and three undescribed species of Brangas . The genus occurs from Mexico to southern Brazil and northern Argentina . Robbins (2004b) placed Thecla teucria Hewitson in Brangas while Bálint & Faynel (2008) moved Brangas teucria to the monotypic genus Dabreras . Brangas teucria possesses a shallow hindwing anal lobe cleft similar to that in Brangas . Males of B. teucria have a male genitalia dorsal vinculum process that lies under and inside of its associated brush organs, as it does in Brangas . Finally, the genitalia of B. teucria and B. rita (Goodson) are almost indistinguishable. Brangas teucria (Hewitson, 1868) ( Thecla ) , type locality: Brazil Distribution and Habitat . The Amazonian Region in wet lowland forest. Brangas rita (Goodson, 1945) ( Thecla ) , type locality: Colombia Distribution and Habitat . Costa Rica to western Colombia and Ecuador in wet lowland forest. Brangas dydimaon (Cramer, 1777) ( Papilio ) , type locality: Surinam Distribution and Habitat . The Amazonian Region in wet lowland forest. Brangas neora (Hewitson, 1867) ( Thecla ) , type locality: Guatemala Distribution and Habitat . Mexico to southern Brazil in wet and dry lowland forest, occasionally occurring in montane habitats. Brangas moserorum Bálint & Faynel, 2008 , type locality: Brazil (SP) Distribution and Habitat . Paraguay and southern Brazil , from MT, DF, and MG to RS. Brangas coccineifrons ( Godman & Salvin, 1887 ) ( Thecla ) , type locality: Nicaragua ; Colombia Distribution and Habitat . Mexico to western Ecuador in wet lower montane forest, at lower elevations in the north. Brangas torfrida (Hewitson, 1867) ( Thecla ) , type locality: Brazil (PA) Distribution and Habitat . The Amazonian and Atlantic regions in wet lowland forest to lower montane forest. Brangas carthaea (Hewitson, 1868) ( Thecla ) , type locality: Mexico Distribution and Habitat . Mexico to western Ecuador in wet montane forest, but at lower elevations in the north. Brangas getus (Fabricius, 1787) ( Papilio ) , type locality: Surinam , repl. name pelops (Stoll, 1781) ( Papilio ), type locality: Surinam , preocc. (not Drury, 1773) Distribution and Habitat . Mexico to Argentina , Paraguay , and southern Brazil in wet and dry lowland forest. Brangas caranus (Stoll, 1780) ( Papilio ) , type locality: Surinam Distribution and Habitat . From western Ecuador to French Guiana , south to southern Brazil (ES) in wet lowland forest. Brangas rambutorum Bálint & Faynel, 2008 , type locality: Colombia Distribution and Habitat . Venezuela and Colombia in dry forest. Brangas neildonatus Bálint & Faynel, 2008 , type locality: Venezuela Distribution and Habitat . Venezuela . Dry lowland forest along the Venezuelan northern coast. Brangas silumena (Hewitson, 1867) ( Thecla ) , type locality:? Distribution and Habitat . Paraguay and Brazil from RJ to RS in montane forest, but at lower elevations in the south. Brangas felderi (Goodson, 1945) ( Thecla ) , type locality: Colombia ; Ecuador contrastus Bálint, 2008 , New Synonym , type locality: Ecuador Distribution and Habitat . Colombia to Peru in montane forest. Synonymy . The differences between B. felderi and B. contrastus mentioned by Bálint (width of the dorsal forewing border, amount of ventral blue iridescence) represent a small part of continuous variation in the series of more than 25 males that we have examined from eastern Ecuador . Brangas polonus Bálint, 2008 , type locality: Peru Distribution and Habitat . Peru to Bolivia in montane forest. Brangas insolitus Bálint & Faynel, 2008 , type locality: Peru Distribution and Habitat . Ecuador to Peru in montane forest.