New species of Nearctic oak gall wasps (Hymenoptera: Cynipidae, Cynipini)
Author
Melika, George
Author
Nicholls, James A.
0000-0002-9325-563X
james.nicholls@csiro.au
Author
Abrahamson, Warren G.
0000-0002-3557-3613
abrahmsn@bucknell.edu
Author
Buss, Eileen A.
eabuss@ufl.edu
Author
Stone, Graham N.
0000-0002-2737-696X
gstone@staffmail.ed.ac.uk
text
Zootaxa
2021
2021-12-23
5084
1
1
131
journal article
2793
10.11646/zootaxa.5084.1.1
bd42fe03-1a35-4f17-b0fe-55b6ec7fdf80
1175-5326
5800716
53B21C11-CA12-480F-8048-1A0601784172
Neuroterus
Hartig, 1840
Neuroterus
has historically been a complex genus. As originally defined, species of this genus (including the
Neuroterus
species described here) have a mesonotum that is thinner and more weakly sclerotized than is typical for other cynipid genera and the transscutal articulation is absent, either partially (in its median section) or entirely. As a result of the fusion of the mesoscutum (which is slightly invaginated medially) and mesoscutellum, the boundary between these structures is not straight but slightly curved towards the mesoscutum; in addition, the mesoscutum is emarginate posterolaterally and slightly elevated above the dorsoaxillar area. In all other Nearctic
Cynipini
genera the transscutal articulation is straight, as is the boundary between the mesoscutum posteriorly and mesoscutellum anteriorly; however, in some apterous Nearctic genera (
Acraspis
,
Biorhiza
Westwood, 1840
,
Phylloteras
Ashmead, 1897
,
Zopheroteras
Ashmead, 1897
) this articulation is present although very reduced (
Melika & Abrahamson 2002
).
Elevated
heterogeneity within this genus was demonstrated by
Kinsey (1923)
when he subdivided
Neuroterus
into six subgenera on the basis of adult morphology, geographic distribution, gall structure, and life cycles, underlining the fact that this group is biologically diverse and unlikely to be monophyletic. Additional Asian taxa were placed in this genus thus further increasing the heterogeneity (
Monzen 1954
;
Abe 2006
), although the divergent subgenus
Pseudoneuroterus
Kinsey, 1923
was raised to full genus status by
Pujade-Villar
et al
. 2004
. Subsequent phylogenetic reconstructions strongly challenged the monophyly of
Neuroterus
(
Stone
et al
. 2009
,
Liljeblad
et al
. 2008
), prompting significant revision of some distinct lineages within the Palaearctic component of
Neuroterus
through creation or re-establishment of the genera
Cerroneuroterus
Melika & Pujade-Villar 2010
,
Trichagalma
Mayr, 1907
and
Latuspina
Monzen, 1954
(
Melika
et al
. 2010
). However, molecular data show that even the most recent definition of
Neuroterus
is still not monophyletic (
Stone
et al
. 2009
;
Nicholls
et al
. 2017
;
Nicholls
et al
. 2018b
;
Nieves-Aldrey
et al
. 2021
).
Currently just over 50 species of
Neuroterus
are known from America north of
Mexico
and a further 15 species from
Mexico
and central America (
Kinsey 1938
;
Burks 1979
;
Lyon 1984
;
Melika & Abrahamson 1997a
; Pujade-Villar
et al
. 2015a, 2016, 2017c, 2018b;
Pujade-Villar & Hanson 2021
;
Medianero & Nieves-Aldrey 2017
). Nearly all of these species induce galls on
Quercus
section
Quercus
with the exception of one species with its host plant in section
Protobalanus
, four species that gall hosts in section
Virentes
and a single species galling the section Ponticae; the latter two oak sections were recently split out from a broader definition of section
Quercus
by
Denk
et al
. (2017)
. Herein we describe seven new species of
Neuroterus
, six of which are associated with
Quercus
section
Quercus
and one with section
Virentes
.