Phylogeny and classification of armored scale insects (Hemiptera: Coccomorpha: Diaspididae)
Author
Normark, Benjamin B.
Author
Okusu, Akiko
Author
Morse, Geoffrey E.
Author
Peterson, Daniel A.
Author
Itioka, Takao
Author
Schneider, Scott A.
text
Zootaxa
2019
2019-06-17
4616
1
1
98
journal article
26473
10.11646/zootaxa.4616.1.1
324b030a-18a2-4dac-9ddd-28c468aea7df
1175-5326
3248018
1B9DFBC9-2BA8-4619-8F70-E372DCBD7411
Tribe
DIASPIDINI Targioni Tozzetti
Diagnosis: AF
without fringed plates (except in some bamboo-feeding species), with gland spines; L2 bilobular; ducts 2-barred; long axes of orifices of marginal macroducts usually oblique or perpendicular to body margin; pores by anterior spiracle usually present, usually 3-locular; marginal macroducts usually the same size as, or only slightly larger than, dorsal macroducts.
2F
similar to AF.
CR
without seta on tarsus.
AM
postoccipital ridge not produced posteriorly into a median process; anterior arms of postoccipital ridge separated by a distance equal to or greater than their own length; with 1 pair of tentorial pits.
2M
usually with tricarinate scale cover and distinctly different from 2F, with more dorsal ducts and often having modified ducts unlike those seen in females.
FIGURE 7.
Phylogeny of the subtribe Diaspidina (Diaspidinae:
Diaspidini
), whose relationship to other
Diaspidini
is shown in Fig. 2. For further explanation of the analysis and notation, see the caption to Fig. 1.
This tribe is notable for the marked sexual dimorphism of the second-instar nymph. Male scale covers in the field often have 3 distinctive ridges or carinae, and on a microscope slide second-instar males often have distinctive ducts not seen in other life stages; these differ between the subtribes as described below.
Included genera:
Furchadaspis
MacGillivray
,
Gadaspis
Borchsenius
,
Ischnaspis
Douglas
,
Nimbaspis
Balachowsky
,
Prodiaspis
Young
,
Pygalataspis
Ferris
,
Roureaspis
Takagi
,
Trischnaspis
Ben-Dov
, and the genera listed in the subtribes below.
Distribution:
Cosmopolitan.