Notes on the genus Anomala SAMOUELLE, 1819 (Coleoptera, Scarabaeidae, Rutelinae) in Vietnam and neighboring regions: eight new species and faunistic records
Author
Zorn, Carsten
Author
Kobayashi, Hirokazu
Author
Wada, Kaoru
text
Beiträge Zur Entomologie = Contributions to Entomology
2017
67
2
325
352
journal article
10.21248/contrib.entomol.67.2.325-352
0005-805X
5742448
F2CE9642-9537-46FF-BA3F-5D4148AC1A0B
Anomala herbacea
spec. nov.
urn:lsid:zoobank.org:act:
D39067BB-B013-4D67-9E01-BEA126CF5EA8
(
Figs 9
,
17
)
Type series
:
Holotype
: , C.
VIETNAM
:
Kon Tum
dist nr.
Ngoe Thinh
MAY. 2010
. (
NSMT
)
.
Paratypes
: 1 , the same data as for the holotype (
MFPC
); 1 , C. VIET- NAM:
Kon Tum Province
, ca.
30 km
from
Kon Plong
,
1250 m
14°40,320N
108°15,829E
| leg.
L. Bartolozzi
, A.
Bandinelli, S
.
Bambi, V
. Sbordoni at light
4-7.V.2016
(n°Mag. 3078) (
MZUF
)
.
Description
: Body shape elongate ovoid, weakly convex; body length:
14.7-15.7 mm
, width: 8.8-9.0 mm.
Color
. Dorsal surface light green with metallic shine, elytra with brown suffusions; pronotum with narrow yellow band along lateral margins; front of clypeus and antennae yellowish brown; ventral surface and legs yellowish brown with weak metallic reflections except for mesotarsi, metatibia and -tarsi (greenish to reddish copper) and abdominal ventrites (reddish brown).
Head
with erect yellowish brown setae (
0.07-0.38 mm
in length) becoming sparser and shorter anteriad, longer and denser latero-posteriad; interocular distance equals 0.65 times the maximum transverse head width. Antennal club 0.9 times as long as footstalk. Clypeus semicircular, finely and densely rugoso-punctate; anterior margin distinctly reflexed; fronto-clypeal suture almost straight. Frons densely and coarsely rugoso-punctate anteriorly and in the middle, punctures becoming sparser and smaller in postero-lateral portions. Vertex coarsely punctate, the punctures somewhat sparser laterad.
Pronotum
1.6 times as wide as long; sides arcuate in the middle, subparallel basad, nearly straight and strongly convergent anteriorly; widest at base; lateral marginal line almost reaching the hind angles; basal marginal line missing; front angles produced and acute, hind angles obtuse, slightly rounded off; surface evenly covered with moderately large, moderately dense, round to weakly transverse punctures; punctures smaller laterad; each puncture bearing an erect yellow seta (
0.15-0.4 mm
in length).
Scutellum
1.6 times as wide as long; apex rounded; punctures like those of disc of pronotum.
Elytra
. Regularly striate-sulcate; intervals weakly convex; each stria marked by a regular line of almost round, hairless punctures; entire surface covered with transverse, seta-bearing (
0.05-0.15 mm
in length) punctures; this irregular punctation sparser on two inner primary costae as well as on the two secondary costae formed within the subsutural interstice; epipleura disappearing shortly before apical curvature which is fringed by thick erect setae; marginal membrane starting near middle of metacoxa.
Pygidium
triangular, widely rounded at apex; weakly tumid; disc with reticulate striolation which is arranged concentrically around the tumidity; with rather dense yellow setosity (
0.1-0.75 mm
in length), setae longer in the apical portion.
Figs 14–17
:
Anomala
spp.
, aedeagus dorsal (
a
), ventral (
b
) and lateral (
c
). –
14
.
A. ordinata
spec. nov.
, holotype.
15
.
A. recordata
spec. nov.,
holotype
.
16
.
A. asaitoae
spec. nov.
,
holotype
.
17
.
A. herbacea
spec. nov.
,
holotype
. Scale =
1 mm
.
Mesoventrite
very narrow and somewhat concave between the approximated mesocoxae.
Metaventrite
with very dense, partly transversely confluent punctures at the sides, sparsely punctate, almost smooth in the middle; with dense, long, pale yellowish brown setae (0.13-1.0 mm in length) at the sides which become sparser and shorter in the middle.
Abdominal ventrites
with rather sparse, transverse punctures (somewhat denser at the sides); each ventrite with a transverse row of semierect yellow setae in the apical half (
0.05-0.13 mm
in length) and denser adpressed setosity laterally; sides of abdominal ventrites 1-4 distinctly carinate.
Legs
slender and long; protibia bidentate; terminal tooth rather short, somewhat obtuse; protarsal segment 5 ventrally with a sharp tooth in middle; inner protarsal claw and outer mesotarsal claw apically incised; the upper branch of the inner protarsal claw small and slender, the lower branch broad, somewhat enlarged ventrally with obtuse tooth in the middle; the upper branch of outer mesotarsal claw slightly more slender than the lower one.
Diagnosis
: This species is easily distinguished from all other known
Anomala
by the combination of a setose dorsal surface, a light metallic green dorsum and sulcate elytra. Furthermore, the shape of the aedeagus with its long, bifid parameres is unique (fig. 17). No described species seems to be closely related. Several species with dorsal setosity occur in
China
and Indo-China and were subsumed in the “
hirsutula
-group” (LIN, 1996).
Anomala herbacea
, however, does not fit in this group because the dorsal setosity is rather semi-erect to erect (adpressed to semi-erect in species of the
hirsutula
-group), the punctures of pronotum and elytra are less transverse, the propygidium is not exposed in
A. herbacea
, and the body is more depressed and broader than in all species of the
hirsutula
-group. In general, the dorsal setosity is a quantitative rather than a qualitative character in
Anomala
, because seta-bearing punctures on pronotum and elytra might be observed in many known species depending on magnification used.
Etymology
: The species epithet is formed by the Latin adjective “herbaceus, -a, -um”. It translates as “grassy” or “grass-colored”. The name refers to the green, hairy dorsal surface of this species.