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<mods:title id="3F98789F1A179C1C6B3D0DE850BDD73D">Cycloachelous levigatus sp. nov., a new swimming crab (Crustacea: Decapoda Portunidae) from the South China Sea</mods:title>
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<taxonomicName id="4C6E5090FFD6FFC0FF56FD5EFE3142DD" authority="Koch, 2021" authorityName="Koch" authorityYear="2021" box="[151,434,690,717]" class="Malacostraca" family="Portunidae" genus="Cycloacheous" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="327" phylum="Arthropoda" rank="species" species="levigatus" status="sp. nov.">
<emphasis id="B91AF701FFD6FFC0FF56FD5EFE3142DD" bold="true" box="[151,434,690,717]" italics="true" pageId="2" pageNumber="327">Cycloacheous levigatus</emphasis>
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(
<figureCitation id="13553796FFD6FFC0FF5EFD3BFE8242E1" box="[159,257,727,753]" captionStart="FIGURE 1" captionStartId="4.[151,250,1955,1979]" captionTargetBox="[200,1379,199,1909]" captionTargetId="figure-32@4.[181,1406,181,1931]" captionTargetPageId="4" captionText="FIGURE 1. Overall habitus.A, Cycloacheous orbitosinus (Rathbun, 1911), male 23 mm (MNHN-IU-2010-6278). B, Cycloachelous levigatus sp. nov., male 23.4 mm (MNHN-IU-2014-10097). Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4766925" httpUri="https://zenodo.org/record/4766925/files/figure.png" pageId="2" pageNumber="327">Figs. 1B</figureCitation>
, 
<figureCitation id="13553796FFD6FFC0FECCFD3BFE9942E1" box="[269,282,727,753]" captionStart="FIGURE 2" captionStartId="5.[151,250,1315,1339]" captionTargetBox="[151,1436,181,1291]" captionTargetId="figure-15@5.[151,1436,181,1291]" captionTargetPageId="5" captionText="FIGURE 2. Comparative morphology, sternal views with details of sexual openings. Cycloacheous orbitosinus (Rathbun, 1911): A, male paratype 33.3 mm (USNM 41084); C, D, female 26.7 mm (MNHN-IU-2010-3290). Cycloachelous levigatus sp. nov.: B, male holotype 23 mm (MNHN-IU-2014-10093); E, F female allotype 15.2 mm (MNHN-IU-2014-10091). Scale bars: A–C, E—5 mm, D, F—1 mm. Photo A, by N. Evans." figureDoi="http://doi.org/10.5281/zenodo.4766927" httpUri="https://zenodo.org/record/4766927/files/figure.png" pageId="2" pageNumber="327">2</figureCitation>
BEF, 3D–F, 4A, 5B, 8)
</paragraph>
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<emphasis id="B91AF701FFD6FFC0FF56FD14FE62431F" box="[151,481,759,783]" italics="true" pageId="2" pageNumber="327">
<taxonomicName id="4C6E5090FFD6FFC0FF56FD14FE5E431F" authorityName="J, C. Fabricius" authorityYear="1798" box="[151,477,759,783]" class="Malacostraca" family="Portunidae" genus="Portunus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="327" phylum="Arthropoda" rank="species" species="gladiator" subGenus="Amphitrite">Portunus (Amphitrite) gladiator</taxonomicName>
.
</emphasis>
—de 
<bibRefCitation id="EFFF56E2FFD6FFC0FDD6FD14FD0B431F" author="Haan, W. de" box="[535,648,759,783]" pageId="2" pageNumber="327" pagination="1823 - 1830" refId="ref6363" refString="Haan, W. de (1833 - 1850) Crustacea. In: Siebold, P. F. von (Ed.) Fauna Japonica sive Descriptio Animalium, Quae in Itinere per Japoniam, Jussu et Auspiciis Superiorum, qui Summum in India Batava Imperium Tenent, Suscepto, Annis 1823 - 1830 Collegit, Noitis, Observationibus et Adumbrationibus Illustravit. Lugduni-Batavorum, Leiden, 243 pp. https: // doi. org / 10.5962 / bhl. title. 124951" type="book chapter" year="1833">Haan 1833</bibRefCitation>
, 65, pl. XVIII (1)
</paragraph>
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<taxonomicName id="4C6E5090FFD6FFC0FF56FCFBFEEC433E" authorityName="M. J. Rathbun" authorityYear="1911" box="[151,367,790,814]" class="Malacostraca" family="Portunidae" genus="Portunus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="327" phylum="Arthropoda" rank="species" species="orbitosinus">Portunus orbitosinus</taxonomicName>
.
</emphasis>
— 
<bibRefCitation id="EFFF56E2FFD6FFC0FE4CFCFAFD0D433E" author="Stephenson, W. &amp; Rees, M." box="[397,654,790,814]" pageId="2" pageNumber="327" pagination="1 - 114" refId="ref8354" refString="Stephenson, W. &amp; Rees, M. (1967) Some portunid crabs from the Pacific and Indian Oceans in the collections of the Smithsonian Institution. Proceedings of the United States National Museum, 120, 1 - 114. https: // doi. org / 10.5479 / si. 00963801.120 - 3556.1" type="journal article" year="1967">Stephenson &amp; Rees 1967</bibRefCitation>
, 31–34, figs. 6a–c, e–g (Form A).
</paragraph>
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<taxonomicName id="4C6E5090FFD6FFC0FF56FCD9FE74435C" box="[151,503,820,844]" class="Malacostraca" family="Portunidae" genus="Neptunus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="327" phylum="Arthropoda" rank="species" species="orbitospisnis" subGenus="Achelous">Neptunus (Achelous) orbitospisnis</taxonomicName>
.
</emphasis>
— 
<bibRefCitation id="EFFF56E2FFD6FFC0FDD4FCD8FD04435D" author="Sakai, T." box="[533,647,820,845]" pageId="2" pageNumber="327" pagination="365 - 741" refId="ref7545" refString="Sakai, T. (1939) Studies on the Crabs of Japan IV. Brachygnatha, Brachyrhyncha. Vol. 3. Yokendo Co., Ltd., Tokyo, 365 - 741." type="book chapter" year="1939">Sakai 1939</bibRefCitation>
, 396, fig. 8, pl 81 (3).
</paragraph>
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<taxonomicName id="4C6E5090FFD6FFC0FF56FCB8FEFF437B" box="[151,380,851,875]" class="Malacostraca" family="Portunidae" genus="Portunus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="327" phylum="Arthropoda" rank="species" species="orbitospinus">Portunus orbitospinus</taxonomicName>
.
</emphasis>
— 
<bibRefCitation id="EFFF56E2FFD6FFC0FE58FCBFFD8F437B" author="Sakai, T." box="[409,524,851,875]" pageId="2" pageNumber="327" refId="ref7581" refString="Sakai, T. (1965) The crabs of Sagami Bay collected by His Majesty the Emperor of Japan. Maruzen, Tokyo, 206 pp." type="book" year="1965">Sakai 1965</bibRefCitation>
, 118, 57 (2)
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<taxonomicName id="4C6E5090FFD6FFC0FF56FC9FFD92439A" ID-CoL="4M2JR" authorityName="M. J. Rathbun" authorityYear="1911" box="[151,529,882,906]" class="Malacostraca" family="Portunidae" genus="Portunus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="327" phylum="Arthropoda" rank="species" species="orbitosinus" subGenus="Cycloacheous">Portunus (Cycloacheous) orbitosinus</taxonomicName>
.—
</emphasis>
<bibRefCitation id="EFFF56E2FFD6FFC0FDEEFC9EFD21439A" author="Sakai, T." box="[559,674,882,906]" pageId="2" pageNumber="327" refId="ref7609" refString="Sakai, T. (1976) Crabs of Japan and the Adjacent Seas. Kodansha, Tokyo, xxxix + 773 pp." type="book" year="1976">Sakai 1976</bibRefCitation>
, 349, figs. 188a, b; 
<bibRefCitation id="EFFF56E2FFD6FFC0FCA8FC9EFBE8439A" author="Yamaguchi, T &amp; Baba, K." box="[873,1131,882,906]" pageId="2" pageNumber="327" pagination="145 - 570" refId="ref8726" refString="Yamaguchi, T, &amp; Baba, K. (1993) Crustacean specimens collected in Japan by Ph. F. von Siebold an H. Burger and held by the National Natuurhistorisch Museum in Leiden and other museums. In: Yamaguchi, T. (Ed.), Ph. F. von Siebold and Natural History of Japan, Crustacea. The Carcinological Society of Japan, Tokyo, pp. 145 - 570." type="book chapter" year="1993">
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&amp; Baba 1993
</bibRefCitation>
, 40–403, fig. 140. 
<emphasis id="B91AF701FFD6FFC0FF56FC7CFE2643B8" box="[151,421,912,936]" italics="true" pageId="2" pageNumber="327">
<taxonomicName id="4C6E5090FFD6FFC0FF56FC7CFE2243B8" baseAuthorityName="Rathbun" baseAuthorityYear="1911" box="[151,417,912,936]" class="Malacostraca" family="Portunidae" genus="Cycloacheous" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="327" phylum="Arthropoda" rank="species" species="orbitosinus">Cycloacheous orbitosinus</taxonomicName>
.
</emphasis>
— 
<bibRefCitation id="EFFF56E2FFD6FFC0FE7FFC7CFD2143B9" author="Spiridonov, V. A. &amp; Neretina, T. V. &amp; Schepetov, D." box="[446,674,912,937]" pageId="2" pageNumber="327" pagination="404 - 429" refId="ref7975" refString="Spiridonov, V. A., Neretina, T. V. &amp; Schepetov, D. (2014) Morphological characterization and molecular phylogeny of Portunoidea Rafinesque, 1815 (Crustacea Brachyura): Implications for understanding evolution of swimming capacity and revision of the family-level classification. Zoologischer Anzeiger, 253, 404 - 429. https: // doi. org / 10.1016 / j. jcz. 2014.03.003" type="journal article" year="2014">
Spiridonov 
<emphasis id="B91AF701FFD6FFC0FDF5FC7DFDEB43B8" box="[564,616,912,936]" italics="true" pageId="2" pageNumber="327">et al.</emphasis>
2014
</bibRefCitation>
, 406, 412.
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