An annotated checklist of the land snail family Orthalicidae (Gastropoda: Pulmonata: Orthalicoidea) in Ecuador, with notes on the distribution of the mainland species
Author
Breure, Abraham S. H.
Author
Borrero, Francisco J.
text
Zootaxa
2008
1768
1
40
journal article
10.5281/zenodo.274265
fcf6c4eb-75e1-45f1-921c-2cbeb8127dc0
1175-5326
274265
Genus
Plekocheilus
Guilding,
1828
Subgenus
Plekocheilus (Plekocheilus)
Guilding,
1828
Type
species:
Voluta aurissileni
Born,
1780
. Distribution: West Indies (
St. Vincent
,
Barbados
),
Venezuela
,
Ecuador
,
Colombia
.
1
.
Plekocheilus (Plekocheilus) cardinalis
(Pfeiffer,
1853
)
Type
locality: Quito.
Lectotype
ZMB
112721
(designated and figured by
Köhler
2007
).
Distribution:?
Napo
(Pilsbry
1899
:
77
);
Pastaza
, Mera (
FLMNH
109841
;
FMNH
86698, 125478
);
Pichincha
, near Mindo, circa
1000 m
(Weyrauch
1967
:
460
); Nanegal; valley of Río Pilatón (both according to Pilsbry
1899
:
77
). “New
Grenada
” (
RMNH
). “
Ecuador
” (
ZMA
).
Remarks: New
Grenada
was used in the
19
th century and is referring to an area comprising
Ecuador
,
Colombia
and parts of
Venezuela
and
Panama
.
|
Subgenus
Plekocheilus (Eurytus)
Albers, 1850
|
|
Type species:
Helix pentadina
d’Orbigny, 1835
.
|
Distribution: |
West |
Indies |
(St. |
Lucia, |
St. |
Vincent), |
|
| Venezuela |
, Brazil, Bolivia, Peru, Ecuador, Colombia.
2
.
Plekocheilus (Eurytus) aristaceus
(Crosse,
1869
)
Type
locality: Quito, reipublicae Aequatoris.
Distribution:
Chimborazo
, Bucay (
IRSN
);
Cotopaxi
, Páramo de Sigchos (
IRSN
);
Pichincha
, near Mindo (
FMNH
216871
)*; Río Pilatón (
IRSN
). “New
Grenada
” (
ZMB
).
Remarks: These are the first precise localities for this species.
3
.
Plekocheilus (Eurytus) aureonitens
(
Miller,
1878
)
Type
locality: Pilatón valley,
1000 m
.
Distribution:
Pichincha
, Río Pilatón. “
Ecuador
” (
ANSP
78577
). Remarks: This species may prove to be a synonym of
Plekocheilus (Eurytus) taylorianus
(Reeve,
1849
)
.
4
.
Plekocheilus (Eurytus) corydon
(Crosse,
1869
)
Type
locality: Quito.
Distribution:
?
[no specific locality known].
5
.
Plekocheilus (Eurytus) doliarius
(Da Costa,
1898
)
Type
locality: Paramba.
Lectotype
BMNH
1907.11.
21.110
(
Breure
1979
; see also
Neubert & Janssen
2004
:
208
, who figure a
paralectotype
(
SMF
9513
) in Pl.
1
Fig.
1
).
Distribution:
Carchi
, Hacienda Paramba (
BMNH
,
IRSN
,
SMF
).
Remarks: As this species is also recorded from
Colombia
, the locality is most likely in Carchi, which is also the only province where Paramba is attributed to in the GNS-gazetteer.
6
.
Plekocheilus (Eurytus) eros
(Angas,
1878
)
Type
locality:
Ecuador
.
Holotype
BMNH
1879.1.21.2. Distribution:
?
[no specific locality known].
7
.
Plekocheilus (Eurytus) floccosus
(Spix,
1827
)
Type
locality:
Brazil
.
Holotype
ZSM
(
Simone
2006
:
149
).
Distribution:
Morona-Santiago
, Cusuimi (
FMNH
170620
); Río Macuma, ca.
10 km
from Río Morona (
FMNH
187606
)*;
Napo
, Río
Napo
,
500 m
(
SMF
156306
; see Weyrauch
1967
:
462
);
Tungurahua
, Topo (
FMNH
86667
); 0.7 km W Río Negro (
RMNH
)*. “
Ecuador
” (
RMNH
).
Remarks: This is a broad ranged species that is also known from
Peru
,
Brazil
and
Bolivia
. Weyrauch (
1967
:
462
) placed
Helix pentadina
d’Orbigny,
1835
in the synonymy of this species.
8
.
Plekocheilus (Eurytus) jimenezi
(Hidalgo,
1872
)
Type
locality: San José (according to Weyrauch
1967
:
463
this is located
50 km
E Baeza; it is most probably San José de Suno [
00°
32
' 00" S
077°
25
' 00" W
], also known as San José Viejo).
Plekocheilus (Eurytus) gibbonius
(Hidalgo,
1872
), described from the same locality, is a synonym according to
Pilsbry (
1895
:
87
)
. Distribution:
Orellana
, San José de Suno. “Oriente Prov.” (
FMNH
72336
).
Plekocheilus (Eurytus) jimenezi oligostylus
Pilsbry,
1939
Type
locality: [
Ecuador
, between Baeza and Archidona, Nachiyacu, see Clench & Turner
1962
:
109
].
Holotype
:
ANSP
170696
.
Distribution:
Napo
, Nachiyacu; Sarayacu (see Weyrauch
1967
:
463
); valley of Río Quijos (
FMNH
216872
);
Pastaza
, Puyo (
FLMNH
109855
;
FMNH
42168, 126934
); Mera (
FLMNH
184702
).
9
.
Plekocheilus (Eurytus) lynciculus
(Deville & Hupé,
1850
)
Type
locality:
Peru
, Loreto, Sarayacu.
Distribution:
Napo
, Nachiyacu (
ANSP
170694
:
Holotype
of the synonym
Plekocheilus (Eurytus) jacksoni
Pilsbry,
1939
; according to Weyrauch
1967
:
463
);
Orellana
, Loreto,
550 m
[
SMF
156302
; see Weyrauch
1967
:
463
];
Pastaza
, Chichirota (
FMNH
78742
).
10
.
Plekocheilus (Eurytus) mcgintyi
‘Pilsbry’ H.B.
Baker
,
1963
Type
locality: Rio
Napo
, northeastern boundary of
Ecuador
.
Holotype
ANSP
227455
. Distribution:
Napo
, Río Jatunyacu [= Río
Napo
] (
ANSP
).
11
.
Plekocheilus (Eurytus) nocturnus
Pilsbry,
1939
Type
locality:
Ecuador
, Puyo.
Holotype
ANSP
170695
. Distribution:
Pastaza
, Puyo (
FMNH
86644
).
12
.
Plekocheilus (Eurytus) phoebus
(Pfeiffer,
1863
)
Type
locality:
Ecuador
.
Lectotype
BMNH
1975143
(
Breure
1979
). Distribution:
?
[no specific locality known].
13
.
Plekocheilus (Eurytus) pulicarius
(Reeve,
1848
)
Type
locality: New
Grenada
.
Lectotype
BMNH
1975281
(designated and figured by
Breure
1978
:
15
, Pl.
11
Fig.
6
).
Distribution:
?
[no specific Ecuadorian locality known; the species is better known in
Colombia
]. “
Ecuador
” (
FMNH
202082
). “New
Grenada
” (
BMNH
1975282
/
2
paralectotypes
).
14
.
Plekocheilus (Eurytus) roseolabrum
(E.A. Smith,
1877
)
Type
locality: Malacatos.
Lectotype
BMNH
1975135
(
Breure
1978
).
Distribution:
Loja
, Malacatos (
BMNH
1877.3.28.2/
1
paralectotype
);
Zamora-Chinchipe
, Tapichalaca* (B. Harris, personal communication and photograph).
Remarks: This is the first record and precise locality after the original description in
1877
.
15
.
Plekocheilus (Eurytus) taylorianus
(Reeve,
1849
)
Type
locality: Environs of Quito.
Lectotype
BMNH
1874.12.
11.271
(Here corrected for
BMNH
1975142
as given by
Breure
1978
).
Pilsbry (
1895
)
placed
Eurytus taylorioides
Miller,
1878
in the synonymy of this species; its
type
locality is: “circa Quito, Chimborazo”.
Distribution:
Pastaza
, Mera (
FMNH
202059
);
Pichincha
, Milpe (
FLMNH
166225
); Mindo, near Rio Blanco (
FLMNH
166226
); Nanegal (
EPN
); Pacto (
EPN
); Pintag (
EPN
); Gualea (
FMNH
61138, 31173
). “
Ecuador
” (
CMC
C
11056
,
C
11057
;
FMNH
100722
;
RMNH
;
ZMA
).
Remarks:
Köhler (
2007
)
gives a figure of a
paralectotype
(
ZMB
112722
).
16
.
Plekocheilus (Eurytus) tricolor
(Pfeiffer,
1853
)
Type
locality: New
Grenada
, Gualea. According to
Pilsbry (
1901
:
131
)
Bulimus semipictus
Hidalgo,
1869
is a junior synonym is, for which the
type
locality is Baeza.
Distribution:
Cotopaxi
, Sigchos (
FMNH
70912
);
Imbabura
, Ibarra (
FMNH
70911
);
Napo
, Baeza;
Orellana
, Ávila (
FMNH
31171
);
Pichincha
, Gualea. “
Ecuador
” (
RMNH
).
Subgenus
Plekocheilus (Aeropictus)
Weyrauch,
1967
Type
species:
Bulimus veranyi
Pfeiffer,
1848
. Distribution:
Venezuela
,
Brazil
,
Ecuador
,
Colombia
.
17
.
Plekocheilus (Aeropictus) tenuissimus
(Weyrauch,
1967
)
Type
locality:
20 km
W Quito, Tandayapa,
2500 m
.
Holotype
IML
3364
. Distribution:
Pichincha
, Tandayapa* (
IML
); near Nanegal (
RMNH
)*.
Plekocheilus
sp.
Distribution:
El Oro
, 13.1 km WNW
Pinas
(
FLMNH
36133
)*;
Napo
, Archidona (
FMNH
31133
), labelled as
P. blainvilleanus loveni
(Pfeiffer)
, which is a Venezuelan species;
Pichincha
,
15 km
NE Las Palmas, Río Faisanes (
FLMNH
22917
)*; 14.3 km NE La Palma, Río Faisanes (
FLMNH
26611
)*.