Revision and cladistic analysis of the Guineo-Congolian spider genus Smeringopina Kraus (Araneae, Pholcidae
Author
Huber, Bernhard A.
text
Zootaxa
2013
3713
1
1
160
journal article
10.11646/zootaxa.3713.1.1
935605ac-a0e6-4963-8264-b4b0ccb8568b
1175-5326
284803
C5F0BC11-92C0-4B30-9DB3-200882AC8950
Smeringopina tebe
new species
Figs. 26
,
133–137
,
167, 177
,
225–239
Type
. ♂
holotype
from
Gabon
, Ogooué-Ivindo, N
Tébé
(
0°02.3’S
,
13°40.9’E
),
550 m
a.s.l., forest along brook,
17.viii.2011
(B.A. & S.R. Huber), in ZFMK (Ar 10201).
Other material examined.
GABON
:
Ogooué-Ivindo
: N
Tébé
, same data as
holotype
,
8♂
3♀
1 juv.
in ZFMK (Ar 10202); same data, 2♀
1 juv.
in pure ethanol, in ZFMK (Gab 164).
Etymology. The name is a noun in apposition, derived from the
type
locality.
Diagnosis. Easily distinguished from congeners by highly distinctive frontal apophyses on male chelicerae (
Figs. 228–229
,
232
) and by pair of (probably functionally corresponding) deep pockets in anterior epigynal plate (
Fig. 230
).
Male (
holotype
). Total body length 3.8, carapace width 1.2. Leg 1: 38.1 (8.8 + 0.4 + 9.1 + 18.0 + 1.8), tibia 2: 5.5, tibia 3: 3.8, tibia 4: 5.7; tibia 1 L/d: 89. Distance PME-PME 125 µm, diameter PME 115 µm, distance PME- ALE 60 µm, distance AME-AME 25 µm, diameter AME 95 µm. Carapace ochre-yellow with brown triangular mark posteriorly and brown lateral margins; ocular area posteriorly brown, clypeus and sternum brown; legs light brown, femora with dark rings subdistally, tibiae with dark rings proximally and subdistally; abdomen ochre-gray with dark pattern dorsally, laterally, and ventrally, ventral dark bands with lateral constriction. Habitus as in
Figs. 133–134
, ocular area slightly elevated, secondary eyes with indistinct ‘pseudo-lenses’; clypeus with pointed and slightly hooked apophysis between AME and rim; deep thoracic pit and pair of shallow furrows diverging behind pit. Chelicerae as in
Figs. 228–229
, with lateral apophyses in very distal position, with distinctive frontal apophyses consisting of anterior lobe with granulate surface and posterior lobe with smooth surface (
Figs. 232– 233
), without modified hairs. Palps as in
Figs. 135–137
; coxa with distinct retrolateral apophysis; trochanter barely modified; femur with large retrolateral apophysis directed toward ventrally, with sclerotized hump at prolateral joint to trochanter, with weakly sclerotized ventral projection distally; prolateral femur-patella joint strongly shifted toward ventrally; tarsus with some longer and slightly stronger hairs dorsally, tarsal organ capsulate and on short stalk (
Fig. 235
); procursus as in
Figs. 225–226
, with complex membranous and sclerotized structures ventrally (
Figs. 234, 236
), without hinge; bulb with simple weakly sclerotized process (
Fig. 227
; sperm duct apparently opens at basis of this process). Legs without spines and curved hairs, with few vertical hairs; retrolateral trichobothrium on tibia 1 at 2%; prolateral trichobothrium present on all tibiae; pseudosegments barely visible. ALS with eight spigots each (
Fig. 237
); gonopore with two epiandrous spigots (
Fig. 238
).
Variation. Some males with indistinct additional (third) wide darker ring on tibiae. Tibia
1 in
8 other males: 8.0–9.3 (mean: 8.7).
Female. In general similar to male; clypeus unmodified. Tibia
1 in
three females: 6.0, 6.3, 6.3. Epigynum anterior plate triangular with pair of deep pockets (
Figs. 167
,
230
); posterior plate laterally with overhanging folds; internal genitalia as in
Figs. 177
,
231
,
239
.
Natural history. Litter-dwelling species, very common in humid litter near a brook at the
type
locality.
Distribution. Known from
type
locality only (
Fig. 114
).