Review of the species of Macrodactylus Dejean (Coleoptera: Scarabaeidae Melolonthinae) from the Central American Nucleus Author Arce-Pérez, Roberto Author Morón, Miguel Ángel text Zootaxa 2020 2020-05-11 4772 3 567 584 journal article 22256 10.11646/zootaxa.4772.3.7 8b277405-a91c-4d0c-b77e-8b1d4820c7cd 1175-5326 3819764 8DE2F40F-0931-4002-97C4-5603E363B7E8 Macrodactylus tibialis Arce-Pérez & Morón, 2005 Figs. 54, 75–77, 84 Material examined. 115 specimens : 67 ♂ , 48 ♀ . Diagnosis . Body length 12.0–12.5 mm; clypeus, antennae, pronotum, elytra, pygidium, apical sternite, femora, and tibiae orange; frons, scutellum, pterothorax, abdomen, and tarsi black; dorsally glabrous (Fig. 54), male with a strong, acute spine near the middle of the inner edge of each protibia; tarsi without ring of white setae; parameres stout, glabrous, evenly curved externally, forming an oval in distal view; apices weakly lanceolate (Figs. 75–76); female similar to male except, protibiae without acute spine; pygidium short, weakly convex; genital plates (Fig. 77). Natural history. This species inhabits humid montane forests ranging from 1,000 to 1,800 m (additional information in Arce-Pérez & Morón 2005 , 2011 ). Geographical distribution . Honduras ( El Paraíso , Olancho ) and Nicaragua ( Jinotega , Matagalpa ) (Fig. 84) .