Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities Author Brazidec, Manuel https://orcid.org/0000-0002-0860-8972 Univ Rennes, CNRS, Geosciences Rennes, UMR 6118, 35000, Rennes, France & Institut de Syste ́ matique, E ́ volution, Biodiversite ́ (ISYEB), Muse ́ um national d'Histoire naturelle, CNRS, Sorbonne Universite ́, EPHE, Universite ́ des Antilles, CP 50, 57 rue Cuvier, F- 75005 Paris, France & State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, China manuel.brazidec@gmail.com Author Vilhelmsen, Lars https://orcid.org/0000-0002-5593-5722 Natural History Museum of Denmark, SCIENCE, University of Copenhagen, Universitetsparken 15, DK- 2100, Denmark Author Boudinot, Brendon E. https://orcid.org/0000-0002-4588-0430 Institut fuer Zoologie und Evolutionsforschung, Friedrich-Schiller-Universitaet Jena, Vor dem Neutor 1, 07743 Jena, Germany & National Museum of Natural History, Smithsonian Institution, 10 th & Constitution Ave. NW, Washington, DC, USA & Senckenberg Gesellschaft fuer Naturforschung und Naturmuseum, Frankfurt am Main, 60325 Germany Author Richter, Adrian https://orcid.org/0000-0001-5627-2302 Institut fuer Zoologie und Evolutionsforschung, Friedrich-Schiller-Universitaet Jena, Vor dem Neutor 1, 07743 Jena, Germany & Biodiversity and Biocomplexity Unit, Okinawa Institute of Science and Technology Graduate University, Onna-son, Okinawa, Japan Author Hammel, Joerg U. https://orcid.org/0000-0002-6744-6811 Institute of Materials Physics, Helmholtz-Zentrum Hereon, Max-Planck-Str. 1, 21502 Geesthacht, Germany Author Perkovsky, Evgeny E. https://orcid.org/0000-0002-7959-4379 Natural History Museum of Denmark, SCIENCE, University of Copenhagen, Universitetsparken 15, DK- 2100, Denmark & I. I. Schmalhausen Institut of Zoology, National Academy of Sciences of Ukraine, B. Khmelnitskogo 15, Kiev 01030, Ukraine Author Fan, Yong Fushun Amber Institute, Fushun 113005, China Author Wang, Zhen Natural History Museum of Denmark, SCIENCE, University of Copenhagen, Universitetsparken 15, DK- 2100, Denmark & College of Life Sciences, Capital Normal University, 105 Xisanhuanbeilu, Haidian District, Beijing 100048, China Author Wu, Qiong Natural History Museum of Denmark, SCIENCE, University of Copenhagen, Universitetsparken 15, DK- 2100, Denmark & College of Life Sciences, Capital Normal University, 105 Xisanhuanbeilu, Haidian District, Beijing 100048, China Author Wang, Bo State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, China Author Perrichot, Vincent https://orcid.org/0000-0002-7973-0430 Univ Rennes, CNRS, Geosciences Rennes, UMR 6118, 35000, Rennes, France text Arthropod Systematics & amp; Phylogeny 2024 2024-03-22 82 151 181 http://dx.doi.org/10.3897/asp.82.e111148 journal article http://dx.doi.org/10.3897/asp.82.e111148 1864-8312-82-151 43AC036E93CC4D79939A07DF54BE1A2D 8D187145BFCA55A1A73960CDE5BD124B 3.2.4. Tribe † Valaaini trib. nov. Type genus. † Valaa Perrichot, 2009. Diagnosis. Head hypognathous; compound eye without setation, without posterior carina; subantennal groove without dorsal carina; mandible with three teeth; occipital carina present, curving towards mandible. Median mesoscutal sulcus crenulate; parapsidal lines absent; axillae not connected at inner angles, separated by two triangular foveae; propodeum carinate. Protibia with stout apical spines; pro-, meso- and metatibiae shorter than combined length of tarsomeres. Fore wing hyaline, with C, Sc+R, A, Rs+M, cu-a, Rs, M and R1 tubular; pterostigma reduced; M not reaching apical margin; Rs apically strongly angled, arched towards stigma; costal, submarginal, marginal and radial+first cubital cells closed. Hind wing with Rs short, barely projecting beyond R1. Metasoma compact, shorter than mesosoma. Genera and species included. † Valaa delclosi Perrichot, 2009 Comments. The tribe † Valaaini trib. nov. is erected based on the results of the phylogenetic analyses, because † Valaa is not retrieved within a clade with other Dinapsini s.s., contrary to what was suggested by Perrichot (2009) .