Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil Author Amorim, Gabriela Dos Santos Programa de Pós-graduação em Biologia Vegetal, Universidade Federal de Pernambuco, Av. Prof. Moraes Rego, 1235 - Cidade Universitária, 50670 - 901, Recife - PE, Brazil. Author Barbosa, Maria Regina De Vasconcellos Departamento de Sistemática e Ecologia, Universidade Federal da Paraíba, Cidade Universitária, Caixa Postal 5065, 58051 - 970, João Pessoa- PB, Brazil. Author Jr, Eduardo Bezerra De Almeida Departamento de Biologia, Universidade Federal do Maranhão, Cidade Universitária Dom Delgado, Av. dos Portugueses, 1966 - Vila Bacanga, 65080 - 805, São Luís - MA. Author Thomas, William Wayt 0000-0002-4996-536X Institute of Systematic Botany, New York Botanical Garden, 2900 Southern Boulevard, Bronx, 10458 - 5126, NY, United States. wthomas @ nybg. org; https: // orcid. org / 0000 - 0002 - 4996 - 536 X wthomas@nybg.org text Phytotaxa 2022 2022-10-11 568 1 27 60 http://dx.doi.org/10.11646/phytotaxa.568.1.3 journal article 10.11646/phytotaxa.568.1.3 87ddb879-1616-4e3b-8817-434b61fb0c60 1179-3163 7184245 5. Eugenia cupulata Amshoff (1942: 160) . Type:— SURINAME . Brokopondo , Brownsberg, C. J . Zaandam 6428 ( holotype : U [0005014]) . Trees or shrubs 5.4– 10 m . Twigs glabrous. Leaves with petioles 8–22 mm long, glabrous, unchanneled; blades elliptic to broadly so, lanceolate, ovate or oblong, 12.9–22.4 × 5–10 cm , glabrous adaxially and glabrous or sparsely pubescent abaxially, the indumentum brownish-red, chartaceous, discolorous, sometimes lustrous adaxially, darker abaxially; glands not seen; base cuneate, rounded or obtuse; apex acute, cuneate, obtuse or cuspidate; midvein adaxially flat; lateral veins prominent abaxially, not arched, 12 to 15 pairs; intra-marginal vein 1–6 mm from the margin, margin flat. Inflorescence axillary, raceme simple, the axis 2.7–4.8 cm , pubescent, the indumentum brownish-red, bracts not seen, caducous before anthesis; pedicels 5–6 mm long, pubescent; bracteoles deltate, 1.5–2 × 1–2 mm , pubescent, persistent in the fruit; flower buds globose, 4–7 × 5 mm , not constricted above ovary; calyx-lobes free, four, orbicular, 1.5–2 × 1.8–3 mm , pubescent, not reflexed, persistent in the fruit; ovary 2-locular. Immature fruits globose, 0.5–0.8 × 0.6–0.9 cm , glabrous or sparsely pubescent, surface smooth. Note:— Eugenia cupulata may be recognized by its adaxially flat midvein; intra-marginal vein 1–6 mm from the margin; a simple raceme with brownish-red pubescence; globose flower buds; and free, pubescent calyx-lobes. Distribution and habitat:— Known from Bolivia , French Guiana , Suriname , Venezuela , central, northeastern and northwestern Brazil ( Acre , Amazonas, Maranhão , Pará and Rondônia states) (POWO 2020, Flora do Brasil 2020 ). In Amazonian Maranhão , E . cupulata was found growing in “terra firme” forest on rocky soil. Phenology:— Flowering in February and March and fruiting in April and June. Specimens examined:— BRAZIL . Maranhão : São Luís , 2°34’38” S , 44°12’04” W , February-March 1939 (fl.), R.L. Fróes 11789 ( NY !) ; 1940 (fr.), R.L. Froes 11864 ( NY !) . Arredores de Estiva , 08 August 1980 (st.), M.G. Silva 5738 ( MG !) . Additional specimens examined:— BRAZIL . Maranhão : Carolina , 7°19’58” S , 47°28’10” W , 12 March 2016 (fr.), A.C. Sevilha 5811 (CEN, image). Lorêto , “Ilha das Balsas” , 140 m , 7°30’ S , 45°3’ W , 01 June 1962 (fr.), G. Eiten 4810 ( NY !). Pará : Santarém , Km 35 da estrada do Palhao , arredores do Acampamento do Igarapé Curupira , 04 September 1969 (fl.), M.B. Silva 2493 with R. Souza ( NY !) .