Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil Author Amorim, Gabriela Dos Santos Programa de Pós-graduação em Biologia Vegetal, Universidade Federal de Pernambuco, Av. Prof. Moraes Rego, 1235 - Cidade Universitária, 50670 - 901, Recife - PE, Brazil. Author Barbosa, Maria Regina De Vasconcellos Departamento de Sistemática e Ecologia, Universidade Federal da Paraíba, Cidade Universitária, Caixa Postal 5065, 58051 - 970, João Pessoa- PB, Brazil. Author Jr, Eduardo Bezerra De Almeida Departamento de Biologia, Universidade Federal do Maranhão, Cidade Universitária Dom Delgado, Av. dos Portugueses, 1966 - Vila Bacanga, 65080 - 805, São Luís - MA. Author Thomas, William Wayt 0000-0002-4996-536X Institute of Systematic Botany, New York Botanical Garden, 2900 Southern Boulevard, Bronx, 10458 - 5126, NY, United States. wthomas @ nybg. org; https: // orcid. org / 0000 - 0002 - 4996 - 536 X wthomas@nybg.org text Phytotaxa 2022 2022-10-11 568 1 27 60 http://dx.doi.org/10.11646/phytotaxa.568.1.3 journal article 163382 10.11646/phytotaxa.568.1.3 87ddb879-1616-4e3b-8817-434b61fb0c60 1179-3163 7184245 4. Eugenia caducibracteata Mazine (in Mazine & Souza 2009: 149 ). Type:— BRAZIL , Pará, Almeirim, Gleba Monte Dourado, floresta secundária de terra firme, J.M. Pires & N.T. Silva 2015 ( holotype : MG). ( Fig. 2 E-F.) Shrubs 2– 17 m . Twigs glabrous, sparsely pubescent when young, the indumentum whitish. Leaves with petioles 3–8 mm long, glabrous or puberulent, channeled or shallowly channeled; blades elliptic to broadly so, lanceolate, ovate or oblong, 5.2–14 × 2.5–6.3 cm , glabrous or sparsely pubescent, the indumentum brownish, chartaceous, concolorous; glands conspicuous; base rounded, cuneate or acute; apex acute or attenuate, rarely acuminate; midvein adaxially sulcate; lateral veins visible, not arched, 8 to 10 pairs; intramarginal vein 1–3 from the margin, margin revolute. Inflorescence axillary or terminal, raceme simple, the axis 4.2–10.4 cm long; bracts not seen, caducous before anthesis; pedicels 2–9 mm long, glabrous; bracteoles ovate to broadly so, or deltate, 1–2 × 1–3 mm , glabrous, persistent in the fruit; flower buds globose, 5 × 5 mm , not constricted above ovary; calyx-lobes free, four, orbicular, 2–3.5 × 3–4 mm , glabrous, not reflexed, persistent in the fruit; ovary 2-locular. Immature fruits oblong or ellipsoid, 1.4–1.7 × 1.1–1.3 cm , glabrous or puberulent, surface smooth. Note:— Eugenia caducibracteata may be recognized by its adaxially sulcate leaf midvein; a simple raceme with a long axis; large flower buds (5 × 5 mm ); and bracts that are caducous before anthesis. Distribution and habitat:— Known from northeastern and northwestern Brazil (Amazonas, Maranhão and Pará states) ( Mazine & Souza 2009 ). E . caducibracteata can be found growing in unflooded tropical moist forest ( Mazine & Souza 2009 ) and in Amazonian Maranhão was found growing in secondary forest. Phenology:— Flowering in February and December and fruiting in August. Specimens examined:— BRAZIL . Maranhão : Açailândia , Estrada BR-222, entre Santa Inês e Açailândia , 300m , 4°45’ S , 45°10’ W , 17 December 1978 (fl.), J. Jangoux 559 ( NY !) . Alto Alegre do Pindaré , Alzilândia , Rio Pindaré , 0- 100 m , 3°45’ S , 46° 05’ W , 11 December 1978 (fl.), J. Jangoux & R.P . Bahia 304 (BHCB, image). Buriticupu , Reserva Florestal da CVRD, 46°24’04” S , 46°24’04” W , 08 February 1996 (fl.), F.H. Muniz B2378 (SP, image). Santa Luzia , Alzilândia , Rio Pindaré , 3°45’ S , 46°05’ W , 11 December 1978 (fr.), J. Jangoux 304 ( NY !) . São Luís , February-March 1939 [fl], R.L. Fróes 11773 ( NY !) . São José de Ribamar , Sítio Aguahy , 2°38’47” S , 44°09’05” W , 03 February 2018 (fl.), G.S. Amorim 448 ( MAR !) ; 03 February 2018 (fl.), G.S. Amorim 447 ( MAR !) ; 26 August 2017 (fr.), G.S. Amorim 373 ( MAR !) . Turiaçu , Palmeirinha , 27 February 1983 (fl.), J. U. Santos 994 ( MG !) .