Review of Glyphidocera Walsingham of Costa Rica (Lepidoptera: Gelechioidea: Glyphidoceridae) Author DAVID ADAMSKI text Zootaxa 2005 858 1 158 journal article 10.5281/zenodo.170793 94a0cedb-15bb-4bfd-955e-77200a536045 1175-5334 170793 511FBF55-10F2-4A00-B15B-CBD34EAB023E Glyphidocera vappae , new species (Figs. 1, 204–206, 286, 355 , Map 37 ) Diagnosis.— Glyphidocera vappae is most similar to G. f o r m a e but differs from the latter by having slightly smaller and more narrowly separate costal furcae of the valva, a shorter apical curnutus of the aedeagal vesica, and absence of cornuti within vesica of aedeagus. Description.— Head: Vertex and frontoclypeus brownish yellow; outer surface of labial palpus dark brown intermixed with brownish­yellow scales to apical margin of segment II, segment III dark brown intermixed with few brownish­yellow scales; inner surface of segment I–II pale brownish­yellow scales intermixed with few dark­brown scales, segment III dark brown intermixed with few pale brownish­yellow scales; scape of antenna brownish yellow or brownish yellow intermixed with few dark­brown scales, flagellum with alternating bands of brownish­yellow and brown scales; male 4 th flagellomere dorsally protuberant and curved, forming a deep concavity opposite slightly excavated flagellomeres 5–6 ; concavity nearly closed by a few setiform scales on apex of protuberance of 4 th flagellomere; proboscis brownish yellow. Thorax: Tegula and mesonotum brownish yellow or brownish yellow intermixed with few dark­brown scales. Legs brownish yellow intermixed with brown scales, apical margins of all segments and tarsomeres brownish yellow. Forewing (Figs. 1, 355 ): Length 6.0 – 7.1 mm (n = 52 ), brownish yellow intermixed with few brownish­orange and few darkbrown scales; cell and surrounding areas paler than marginal areas; cell with 2 dark­brown spots, 1 spot near middle, 1 spot near distal end; middle spot large or small, present or absent; distal spot small; a small spot on CuP basiposterior to midcell spot present or absent; margin brownish yellow or demarcated with dark­brown scales. Undersurface brown intermixed with brownish­yellow scales along costal margin; venation with R 3 , R 4 , and R 5 approximate to dorsoanterior part of cell; M 2 and M 3 approximate to dorsoanterior part of cell; M 1 absent; CuA 1 and CuA 2 branched distad of Cubitus beyond cell, extending in a slight curve to margin; CuP slightly evident basally. Hindwing (Fig. 1 ): Brownish gray; venation with Rs and M 1 stalked about 1 / 3 length beyond end of cell; M 2 closer to M 3 than to M 1 ; M 3 branched with CuA 1 beyond cell, about 1 / 4 distance as Rs and M 1 ; CuA 2 arising from cell about 2 / 3 length of Cubitus; 1 A and 2 A separate basally, forming an elliptical support. Abdomen (Fig. 204 ): Male with 3 transverse, irregular rows of sex scales on intersegmental membrane between terga 2–3 . Male Genitalia (Figs. 205–206 ): Uncus elongate, wide at base, broadly constricted near 1 / 3 , narrowly parallel­sided apically, apex narrowly rounded and setose; gnathos sparsely setose, protuberant, wide throughout length, apically upturned; tegumen slightly widened basally; dorsal strut of tegumen with arms convergent, fusing medially from posteroventral margin, extending to base of uncus; costa of valva nearly straight from shallow depression beyond basal, digitate process to dorsal furca; apex widely bifurcate, margin between furcae U­shaped; dorsal and ventral furcae large, divergent, equal in size, greatly extended beyond outer margin of cucullus; sacculus apically twisted, constricting valva slightly before midlength, widening distoventrally, forming an elongate, broadly rounded cucullus; cucullus membranous and setose; vesica of aedeagus sparsely denticulate, bearing a large crescent­shaped, apical cornutus; cornutus with one side slightly shortened, exposing a hollow, basal cavity. Female Genitalia (Fig. 286 ): Papillae anales elongate, wider basally than apically; apophysis posterioris and apophysis anterioris about equal in length; eighth sternum and eighth tergum about equal in width; eighth sternum with posterior margin broadly rounded, anterior margin broadly emarginate; eighth tergum with posterior margin nearly straight, anterior margin broadly emarginate; antrum enlarged posteriorly from a constricted anterior end, posterior margin slightly rounded, posterolateral margin angular; anterior end with a pair of elongate, spinelike processes inside cavity formed from outer wall; ductus bursae short; corpus bursae slightly enlarged anteriorly, sparsely denticulate; ductus seminalis short, about 5 ­coiled, each coil decreasing in size distally; ductus seminalis with a short, 4 ­coiled internal support; side opposite ductus bursae slightly swollen, with a large, internal support. Holotype , ɗ, “Sector Cerro Cocori, F[in]ca de E. Rojas, 150 m , Prov[incia] Limón, COSTA RICA , E. Rojas, 31 Ene. – 21 Feb. 1992 , L­N­ 286000 , 567500 ”, “ INBio : COSTA RICA : CRI 000, 785137 ” [bar code label], “ INBio , ɗ Genitalia Slide by D. Adsamski, No. 470 ” [yellow label], “ PARATYPE , Glyphidocera guaroa , Adamski ” [blue label]. Paratypes ( 51 ɗ): 23 ɗ, Same label data as holotype except, “ CRI 000, 785313 ”, Slide No. 1033 ”; “ CRI 000, 775096 ”, “Slide No. 1034 ”; “ Ene. 1993 ”, “ CRI 001, 402859 ”, “Slide No. 1035 ”; “ Feb. 1993 ”, “ CRI 000, 998790 ”, “Slide No. 425 ”; “ Mar. 1992 ”, “ CRI 000, 363593 ”, “Slide No. 1039 ”; “ CRI 000, 363629 ”, “Slide No. 1040 ”; “ 26 Jun. a 16 Jul. 1992 ”, “ CRI 000, 703169 ”, “Slide No. 1047 ”; “ Jul. 1993 , # 2252 ”, “ CRI 001, 699131 ”, “Slide No. 472 ”; “ Abr. 1992 ”, “ CRI 000, 785511 ”, “Slide No. 1043 ”; “ CRI 000, 785518 ”, “Slide No. 1044 ”; “ CRI 000, 786302 ”, “Slide No. 1045 ”; “ CRI 000, 785544 ”, “Slide No. 1042 ”; “ CRI 000, 786366 ”, “Wing Slide No. 429 ”; “ Set. 1993 ”, “ CRI 001, 141370 ”, “Slide No. 1049 ”; “Oct”, “ CRI 000, 819686 ”, “Slide No. 1050 ”; “ 9 a 30 Nov. ”, “ CRI 000, 931463 ”, “Wing Slide No. 491 ”; “ CRI 000, 931418 ”, “Slide No. 1059 ”; “ Nov. 1991 ”, “ CRI 000, 460144 ”, “Slide No. 1054 ”; “ CRI 000, 501470 ”, “Slide No. 1051 ”; “ CRI 000, 501449 ”, “Slide No. 1052 ”; “ CRI 000, 460169 ”, “Slide No. 1060 ”; “ Dic. 1991 ”, “ CRI 000, 361205 ”, “Slide No. 1055 ”; “ CRI 000, 361269 ”, “Slide No. 1058 ”; 13 ɗ, “Est. Magsasay, P. N. Braulio Carrillo, 200 m , Prov. Here., COSTA RICA , R. Aguilar, Abr. 1991 , L­N­ 264600 , 531100 ”, “ CRI 000, 275099 ”, “ PARATYPE , Glyphidocera guaroa Adamski ”, “Slide No. 1066 ”; “ May 1991 , M.A. Zumbado”, “ CRI 001, 300398 ”, “Slide No. 1073 ”; “ Oct 1990 ”, “ CRI 000, 180818 ”, “Slide No. 1068 ”; “ CRI 000, 180807 ”, “Slide No. 1069 ”; “A. Fernandez, Oct. 1990 ”, “ CRI 000, 179675 ”, “Slide No. 1070 ”; “R. Aguilar, Dic. 1990 ”, “ CRI 000, 701559 ”, “Slide No. 1072 ”; “ Nov. 1990 ”, “ CRI 000, 226705 ”, “Slide No. 1023 ”; “ CRI 000, 226693 ”, “Slide No. 1064 ”; “ CRI 000, 226773 ”, “Slide No. 1024 ”; “ CRI 000, 226735 ”, “Slide No. 1035 ”; “ CRI 000, 226721 ”, “Slide No. 1062 ”; “ CRI 000, 226729 ”, “Slide No. 1063 ”; “ Nov. 1990 , A. Fernandez”, “ CRI 000, 453154 ”, “Slide No. 1065 ”; 4 ɗ, Cerro Tortuguero, 0–120 m , P. N. Tortuguero, Prov. Limón, COSTA RICA , Jul. 1992 , R. Delgado, L­N­ 285000 , 588000 ”, “ CRI 000, 735815 ”, “Slide No. 602 ”; “ Ene. 1993 ”, “ CRI 001, 352695 ”, “Slide No. 597 ”; “Mar.”, “ CRI 001, 353223 ”, “Slide No. 587 ”; “ Mar. 1991 , J. Solano”, “ CRI 000, 197452 ”, “Slide No. 599 ”; 3 ɗ, “Est. Pitilla, 700 m , 9 km , S. Sta. Cecilia, P. N. Guanacaste, Prov. Guan., COSTA RICA , 19 May – 3 Jun. 1993 , P. Rios, L­N­ 330200 , 380200 ”, “ CRI 001, 353855 ”, “Slide No. 773 ”; “P. Rios & C. Moraga, Oct. 1990 ”, “ CRI 000, 197960 ”, “ PARATYPE , Glyphidocera guaroa Adamski ”, “Slide No. 1074 ”; “ 19 – 23 Jun. 1993 , # 2175 ”, “ CRI 001, 835123 ”, “Slide No. 479 ”; 2 Ψ, “P. N. Manuel Antonio, 80 m , Quepos, Prov. Punt., COSTA RICA , Mar. 1993 , G. Varela, L­S­ 370900 , 448800 ”, “ CRI 001, 409698 ”, “Ψ Slide No. 427 ”, “ PARATYPE , Glyphidocera guaroa Adamski ”; “ Jul. 1992 ”, “ CRI 000, 950374 ”, “Ψ Slide No. 426 ”; 1 ɗ, “Est. Esquinas, Pen. de Osa, Prov. Punta., COSTA RICA , 200 m , Set. 1993 , M. Segura, L­S­ 301400 , 542200 , # 2372 ”, “ CRI 001, 661959 ”, “Slide No. 526 ”, “ PARATYPE , Glyphidocera guaroa Adamski ”; 1 ɗ, “Monumento Nacional Guayabo, Prov. Carta., COSTA RICA , 1100 m , 28 Set. – 21 Nov. 1994 , G. Fonseca, L­N­ 217400 , 570000 , # 3285 ”, “ CRI 002, 024721”; 1 ɗ, “Fca. San Gabriel, 2 km , SW Dos Rios, 600 m , Prov. Alaj., COSTA RICA , I Curso Microlepidoptera, Jul. 1990 , L­N­ 318800 , 383500 ”, “ CRI 000, 475377 ”, “Slide No. 522 ”; 1 ɗ, “Est. Queb. Bonita, 50 m , Res. Biol. Carara, Prov. Punt., COSTA RICA , R. Zuniga, Ene. 1991 , L­N­194500, 469850”, “ CRI 000, 580522 ”, “Slide No. 609 ”; 1 ɗ, “ COSTA RICA , Prov. Limón, Sector Cedrales de la Rita, 3 km N del Puente Rio Suerte, Ruta Puerto Lindo, 10 m , Jul. 1996 , E. Rojas, L­N­ 278600 , 566500 , # 7900 ”, “ CRI 002, 454558 ”, “Slide No. 846 ”; 1 ɗ, “ COSTA RICA , Heredia, Est. Biol. La Selva, 50–150 m , 10 ° 26 'N , 84 °01'W , June 1994 , INBio­OET”, “ 29 Junio 1994 , Bosque primario, L/01/ 109 ”, “ CRI 001, 243844 ”, “Slide No. 918 ”, ( 36 in INBio , 15 in USNM ). Distribution ( Map 37 ): Glyphidocera vappae appears cosmopolitan in Costa Rica and likely is distributed more widely throughout Central America and possibly into South America . MAP37. Distributionof Glyphidoceravappae ().Outlyingcirclesindicateadjacentmultiplelocalities. Etymology: The species epithet vappae is derived from the Latin “vappa,” meaning sour vine.