Taxonomic and Zoogeographic Study of the Japanese Phygadeuontinae (Hymenoptera, Ichneumonidae), with Descriptions of 17 New Species Author Watanabe, Kyohei Kanagawa Prefectural Museum of Natural History, 499 Iryuda, Odawara, Kanagawa 250 – 0031, Japan watanabe-k@nh‛kanagawa-museum‛jp text Bulletin of the Kanagawa Prefectural Museum (Natural Science) 2021 2021-03-30 50 55 136 https://doi.org/10.32225/bkpmnh.2021.50_55 journal article 300009 10.32225/bkpmnh.2021.50_55 70224806-ca38-47ca-bd8a-63da3ac532f9 2189-6720 12774639 DD695C62-DC2F-4D32-8D33-52648E28661D Paraphylax politus sp. nov. (New SJN: Izu-minoga-togari-himebachi) ( Figs 24A–F , 65G , 66F ) Etymology . The specific name is from the strongly polished mesoscutum and metasomal tergites. Type series . Holotype : KPM-NK 55079 , F, JAPAN , Tokyo , Izuoshima Is. , Oshima Town , Motomachi , Mt. Omaruyama , Tsubakinomori , 5–31. X. 2012 , K. Tsujii leg. (MsT). Description . Female (n=1). Body length 4.0 mm. Body polished, covered with silver setae. Head. Clypeus 2.0 times as wide as maximum length, its anterior margin rounded. Face densely punctate, with a median weak convexity ( Fig. 24B ). MSL 0.6 times as long as BWM. Frons, gena, and vertex finely and sparsely punctate. OOL slightly longer than POL and OD. POL as long as OD. Occipital carina complete, its lower end joined with hypostomal carina distant from mandibular base. Upper tooth of mandible as long as lower tooth. Base of mandible flat. Antenna with 21 flagellomeres. Subapical part of flagellum slightly wider than basal part. FL III 5.0 times as long as maximum depth in lateral view. Mesosoma. Upper side of collar without a median carina. Lateral part of pronotum smooth except for narrowly punctate dorsally and ventrally. Epomia long ( Fig. 24D ). Mesoscutum smooth, with a few, sparse, and fine punctures and a median longitudinal groove on the median lobe ( Fig. 24D ). Notaulus long, its posterior ends closed each other ( Fig. 24D ). Scutellum finely and sparsely punctate, with a lateral longitudinal carina except for apical half. Mesopleuron smooth except for areas below subtegular ridge and speculum and along sternaulus finely and densely punctate. Posterior transverse carina of mesosternum complete. Metapleuron punctate, with a complete juxtacoxal carina. Anterior and posterior transverse carinae of propodeum complete ( Fig. 65G ). Pleural carina complete. Lateral longitudinal carina of propodeum present except for anterior section. Area basalis distinct ( Fig. 65G ). Area superomedia distinct ( Fig. 65G ). Propodeum largely smooth, with fine punctures. Length of fore wing 3.2 mm . Areolet absent ( Fig. 24E ). Vein 2m-cu of fore wing with two bullae. Nervellus inclivous, intercepted slightly behind middle. Hind femur 4.1 times as long as maximum depth in lateral view. Hind TS I: II: III: IV: V = 2.0: 0.9: 0.55: 0.3: 0.5. Tarsal claws simple. Metasoma. T I 1.75 times as long as maximum width, smooth. partly coriaceous. Median dorsal carina of T I present except for posterior part. Dorsolateral carina of T I complete. T II to T V smooth ( Figs 24C, F ). Ovipositor sheath 0.95 times as long as hind tibia. Ovipositor straight, with a weak nodus and ventral teeth ( Fig. 66F ). Coloration ( Figs 24A–F ). Body (excluding wings and legs) black to blackish brown. Basal part of antenna yellowish brown. Apex of FL IV to base of FL XI white dorsally. Mandible yellow except for darkened apex. Clypeus and malar space more or less tinged with reddish brown. Palpi white. Pronotum reddish yellow. Tegula yellow. Subtegular ridge reddish yellow. Mesopleuron tinged with dark reddish brown medially. Metasomal tergites dark brown to blackish brown except for posterior parts of T I and T II yellowish brown. Membranous part of metasomal sternites whitish yellow. Ovipositor reddish brown. Wings hyaline, with a longitudinal clouded band just behind pterostigma and a small, clouded area subbasally. Veins yellowish brown to blackish brown. Pterostigma blackish brown. Legs reddish brown. Coxae except for dorsal surface of hind coxa, trochanters, trochantelli except for apical part of hind trochantellus, and tibial spurs whitish yellow. Dorsal surface of hind coxa, hind femur, and hind tibia except for base more or less darkened. Base of hind tibia white. Male. Unknown. Distribution . Japan (Izuoshima Is.). Bionomics . Unknown. Remarks . This species resembles Pa. sakabei Momoi, 1966 in the smooth T III but it can be distinguished by the above key.