<document id="9D009246A3136D503DADFF9B70EF5791" ID="10.11646/zootaxa.4140.1.1" ID-DOI="10.11646/zootaxa.4140.1.1" ID-GBIF-Dataset="01995ca4-7d04-4402-b21a-926957b1a391" ID-ISSN="1175-5326" ID-Zenodo-Dep="256507" ID-ZooBank="AD1F6B4E-3B0D-422C-B47C-0495BD385857" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1469426945103" checkinUser="plazi" docAuthor="Nguyen, Tung T., Nguyen, Anh D., Tran, Binh T. T. &amp; Blakemore, Robert J." docDate="2016" docId="B5727931FF98792EFF3C25D661678E4C" docLanguage="en" docName="zootaxa.4140.1.1.pdf" docOrigin="Zootaxa 4140 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Amynthas papulosus subsp. savanakhetanus" docType="treatment" docVersion="22" lastPageNumber="42" masterDocId="494B0149FFB17907FFAB254162078C42" masterDocTitle="A comprehensive checklist of earthworm species and subspecies from Vietnam (Annelida: Clitellata: Oligochaeta: Almidae, Eudrilidae, Glossoscolecidae, Lumbricidae, Megascolecidae, Moniligastridae, Ocnerodrilidae, Octochaetidae)" masterLastPageNumber="92" masterPageNumber="1" pageNumber="42" updateTime="1698365898991" updateUser="plazi">
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<mods:title id="C10B975144D377DE1E604609AFBB65E8">A comprehensive checklist of earthworm species and subspecies from Vietnam (Annelida: Clitellata: Oligochaeta: Almidae, Eudrilidae, Glossoscolecidae, Lumbricidae, Megascolecidae, Moniligastridae, Ocnerodrilidae, Octochaetidae)</mods:title>
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<mods:namePart id="1C6654D94ADBA0D711355F84B0BA3EF7">Nguyen, Tung T.</mods:namePart>
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<mods:namePart id="527D5A1228224F1411B01B4C12AB2590">Nguyen, Anh D.</mods:namePart>
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<mods:namePart id="134830D1BF2D563E0E2DE8689A029408">Tran, Binh T. T.</mods:namePart>
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<mods:namePart id="12A81EBC2A7C3A4A3DB1D77702F4C6A0">Blakemore, Robert J.</mods:namePart>
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<treatment id="B5727931FF98792EFF3C25D661678E4C" ID-DOI="http://doi.org/10.5281/zenodo.5467216" ID-GBIF-Taxon="123207304" ID-Zenodo-Dep="5467216" LSID="urn:lsid:plazi:treatment:B5727931FF98792EFF3C25D661678E4C" httpUri="http://treatment.plazi.org/id/B5727931FF98792EFF3C25D661678E4C" lastPageNumber="42" pageId="41" pageNumber="42">
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<paragraph id="3D64C827FF98792EFF3C25D661708CF2" blockId="41.[151,887,151,176]" box="[151,887,151,176]" pageId="41" pageNumber="42">
<heading id="662C7F4BFF98792EFF3C25D661708CF2" bold="true" box="[151,887,151,176]" fontSize="10" level="2" pageId="41" pageNumber="42" reason="2">
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<taxonomicName id="FADBB3A4FF98792EFF3C25D6604F8CF2" box="[151,584,151,176]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="41" pageNumber="42" phylum="Annelida" rank="subSpecies" species="papulosus" subSpecies="savanakhetanus">
<emphasis id="0FAF1435FF98792EFF3C25D6604F8CF2" bold="true" box="[151,584,151,176]" italics="true" pageId="41" pageNumber="42">Amynthas papulosus savanakhetanus</emphasis>
</taxonomicName>
(
<bibRefCitation id="594AB5D6FF98792EFDFC25D661688CF2" author="Thai" box="[599,879,151,176]" pageId="41" pageNumber="42" refString="Thai, T. B. &amp; Samphon, K. (1988) New subspecies and species of earthworms (Oligochaeta) from Laos PDR. Journal of Science of HNUE, special issue, 3 - 25." type="book chapter" year="1988">Thai &amp; Samphon, 1988</bibRefCitation>
)
</emphasis>
</heading>
</paragraph>
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<subSubSection id="75C19BACFF98792EFF3C25A161CE8D56" pageId="41" pageNumber="42" type="reference_group">
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<taxonomicName id="FADBB3A4FF98792EFF3C25A160068CB4" box="[151,513,224,246]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="41" pageNumber="42" phylum="Annelida" rank="subSpecies" species="papulosa" subSpecies="savanakhetana">
<emphasis id="0FAF1435FF98792EFF3C25A160068CB4" box="[151,513,224,246]" italics="true" pageId="41" pageNumber="42">Pheretima papulosa savanakhetana</emphasis>
</taxonomicName>
<treatmentCitation id="BC7AEE36FF98792EFDAC25A161198CB4" author="Thai" box="[519,798,224,246]" page="21" pageId="41" pageNumber="42" year="1988">
<bibRefCitation id="594AB5D6FF98792EFDAC25A160FF8CB4" author="Thai" box="[519,760,224,246]" pageId="41" pageNumber="42" refString="Thai, T. B. &amp; Samphon, K. (1988) New subspecies and species of earthworms (Oligochaeta) from Laos PDR. Journal of Science of HNUE, special issue, 3 - 25." type="book chapter" year="1988">Thai &amp; Samphon, 1988</bibRefCitation>
: 21
</treatmentCitation>
; 
<treatmentCitation id="BC7AEE36FF98792EFC8225A061B68CB4" author="Le" box="[809,945,224,246]" page="38" pageId="41" pageNumber="42" year="1995">
<bibRefCitation id="594AB5D6FF98792EFC8225A0618C8CB4" author="Le" box="[809,907,224,246]" pageId="41" pageNumber="42" refString="Le, V. T. (1995 a) The earthworm fauna of Northeastern Vietnam. PhD dissertation in Zoology. Hanoi National University of Education, Hanoi, 148 pp." type="book" year="1995" yearSuffix="a">Le 1995a</bibRefCitation>
: 38
</treatmentCitation>
; 
<treatmentCitation id="BC7AEE36FF98792EFC1025A166638CB4" author="Thai" box="[955,1124,224,246]" page="309" pageId="41" pageNumber="42" year="2000">
<bibRefCitation id="594AB5D6FF98792EFC1025A166358CB4" author="Thai" box="[955,1074,224,246]" pageId="41" pageNumber="42" refString="Thai, T. B. (2000 a) Species diversity of earthworms in Vietnam. Proceedings of the national workshop on the basic issues in life science. Hanoi Science and Technics Publishing House, 307 - 311." type="proceedings paper" year="2000" yearSuffix="a">Thai 2000a</bibRefCitation>
: 309
</treatmentCitation>
. 
<emphasis id="0FAF1435FF98792EFF3C25BF60218D56" box="[151,550,254,276]" italics="true" pageId="41" pageNumber="42">
<taxonomicName id="FADBB3A4FF98792EFF3C25BF636F8D56" box="[151,360,254,276]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="41" pageNumber="42" phylum="Annelida" rank="species" species="papulosus">Amynthas papulosus</taxonomicName>
savanakhetanus—
</emphasis>
<treatmentCitation id="BC7AEE36FF98792EFD8D25BF61048D56" author="Blakemore" box="[550,771,254,276]" page="76" pageId="41" pageNumber="42" year="2007">
<bibRefCitation id="594AB5D6FF98792EFD8D25BF60DA8D56" author="Blakemore" box="[550,733,254,276]" pageId="41" pageNumber="42" refString="Blakemore, R. J. (2007 a) Updated checklist of Pheretimoids (Oligochaeta: Megascolecidae: Pheretima auct.) taxa. www. annelida. net / earthworm / Pheretimoids. pdf (accessed 6 Jun 2016)" type="journal article" year="2007" yearSuffix="a">Blakemore 2007a</bibRefCitation>
: 76
</treatmentCitation>
; 
<bibRefCitation id="594AB5D6FF98792EFCA625BF61C28D56" author="Blakemore" box="[781,965,254,276]" pageId="41" pageNumber="42" refString="Blakemore, R. J. (2008 b) List of earthworms from Cambodia, Laos, and Vietnam (Excel spreadsheet). In: A Series of Searchable Texts on Earthworm Biodiversity, Ecology and Systematics from Various Regions of the World - 2 nd Edition (2008). VermEcology, Yokohama. CD - ROM. Avaliable from: http: // www. annelida. net / earthworm / Vietnam % 20 + % 20 Lao s % 20 + % 20 Cambodia 1. xls (Accessed 21 Jul. 2016)" type="journal article" year="2008" yearSuffix="b">Blakemore 2008b</bibRefCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="75C19BACFF98792EFF3C2400618B8DE0" pageId="41" pageNumber="42" type="materials_examined">
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<materialsCitation id="8DB3C27AFF98792EFF3C240060128D18" ID-GBIF-Occurrence="1288043167" box="[151,533,321,346]" country="Laos" location="Laos" pageId="41" pageNumber="42" specimenCount="1" typeStatus="holotype">
<emphasis id="0FAF1435FF98792EFF3C240063328D18" bold="true" box="[151,309,321,346]" pageId="41" pageNumber="42">
<typeStatus id="E2607685FF98792EFF3C240062D58D18" box="[151,210,321,346]" pageId="41" pageNumber="42">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="45CC88B7FF98792EFE97240363758D1B" box="[316,370,322,345]" name="Laos" pageId="41" pageNumber="42">Laos</collectingCountry>
(Savannaket).
</materialsCitation>
</paragraph>
<paragraph id="3D64C827FF98792EFF6C2424603E8D3C" blockId="41.[151,1436,321,526]" box="[199,569,357,382]" pageId="41" pageNumber="42">
<materialsCitation id="8DB3C27AFF98792EFF6C2424603E8D3C" ID-GBIF-Occurrence="1288043142" box="[199,569,357,382]" collectionCode="SORC" country="Vietnam" location="Vietnam" pageId="41" pageNumber="42" specimenCount="1" typeStatus="holotype">
<emphasis id="0FAF1435FF98792EFF6C242463728D3C" bold="true" box="[199,373,357,382]" pageId="41" pageNumber="42">
<typeStatus id="E2607685FF98792EFF6C242463058D3C" box="[199,258,357,382]" pageId="41" pageNumber="42">Type</typeStatus>
material.
</emphasis>
<collectionCode id="5BCA50E2FF98792EFED7242763CF8D3C" box="[380,456,358,382]" pageId="41" pageNumber="42">SORC</collectionCode>
, 
<collectingCountry id="45CC88B7FF98792EFE78242460318D3C" box="[467,566,357,382]" name="Vietnam" pageId="41" pageNumber="42">Vietnam</collectingCountry>
.
</materialsCitation>
</paragraph>
<paragraph id="3D64C827FF98792EFF6C24C8618B8DE0" blockId="41.[151,1436,321,526]" box="[199,908,393,418]" pageId="41" pageNumber="42">
<materialsCitation id="8DB3C27AFF98792EFF6C24C8618B8DE0" ID-GBIF-Occurrence="1288043174" box="[199,908,393,418]" location="Vietnam. Quang Ninh" pageId="41" pageNumber="42" specimenCount="1" stateProvince="Vietnam. Quang Ninh">
<emphasis id="0FAF1435FF98792EFF6C24C8607F8DE0" bold="true" box="[199,632,393,418]" pageId="41" pageNumber="42">
Records from 
<collectingRegion id="FF1F06C5FF98792EFEDA24C8607F8DE0" box="[369,632,393,418]" inferredBySuffix="true" pageId="41" pageNumber="42">
Vietnam. 
<collectingRegion id="FF1F06C5FF98792EFE4E24C8607F8DE0" box="[485,632,393,418]" country="Vietnam" name="Quang Ninh" pageId="41" pageNumber="42">Quang Ninh</collectingRegion>
</collectingRegion>
</emphasis>
(Mong Cai) (
<bibRefCitation id="594AB5D6FF98792EFCBA24C861878DE0" author="Le" box="[785,896,393,418]" pageId="41" pageNumber="42" refString="Le, V. T. (1995 a) The earthworm fauna of Northeastern Vietnam. PhD dissertation in Zoology. Hanoi National University of Education, Hanoi, 148 pp." type="book" year="1995" yearSuffix="a">Le 1995a</bibRefCitation>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="75C19BACFF98792EFF6C24EC60608D84" box="[199,615,429,454]" pageId="41" pageNumber="42" type="distribution">
<paragraph id="3D64C827FF98792EFF6C24EC60608D84" blockId="41.[151,1436,321,526]" box="[199,615,429,454]" pageId="41" pageNumber="42">
<emphasis id="0FAF1435FF98792EFF6C24EC63678D84" bold="true" box="[199,352,429,454]" pageId="41" pageNumber="42">Distribution.</emphasis>
<collectingCountry id="45CC88B7FF98792EFECC24EF639A8D87" box="[359,413,430,453]" name="Laos" pageId="41" pageNumber="42">Laos</collectingCountry>
(
<bibRefCitation id="594AB5D6FF98792EFE0724EC605D8D84" author="Samphon" box="[428,602,429,454]" pageId="41" pageNumber="42" refString="Samphon, K. (1990) The earthworm fauna of Laos PDR. PhD dissertation. Hanoi National University of Education." type="book" year="1990">Samphon 1990</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="75C19BACFF98792EFF6C249061678E4C" pageId="41" pageNumber="42" type="discussion">
<paragraph id="3D64C827FF98792EFF6C249061678E4C" blockId="41.[151,1436,321,526]" pageId="41" pageNumber="42">
<emphasis id="0FAF1435FF98792EFF6C2490633C8DA8" bold="true" box="[199,315,465,490]" pageId="41" pageNumber="42">Remarks.</emphasis>
<bibRefCitation id="594AB5D6FF98792EFEED2490601D8DAB" author="Blakemore" box="[326,538,465,490]" pageId="41" pageNumber="42" refString="Blakemore, R. J. (2007 a) Updated checklist of Pheretimoids (Oligochaeta: Megascolecidae: Pheretima auct.) taxa. www. annelida. net / earthworm / Pheretimoids. pdf (accessed 6 Jun 2016)" type="journal article" year="2007" yearSuffix="a">Blakemore (2007a</bibRefCitation>
, 
<bibRefCitation id="594AB5D6FF98792EFD86249060748DA8" author="Blakemore" box="[557,627,465,490]" pageId="41" pageNumber="42" refString="Blakemore, R. J. (2008 b) List of earthworms from Cambodia, Laos, and Vietnam (Excel spreadsheet). In: A Series of Searchable Texts on Earthworm Biodiversity, Ecology and Systematics from Various Regions of the World - 2 nd Edition (2008). VermEcology, Yokohama. CD - ROM. Avaliable from: http: // www. annelida. net / earthworm / Vietnam % 20 + % 20 Lao s % 20 + % 20 Cambodia 1. xls (Accessed 21 Jul. 2016)" type="journal article" year="2008" yearSuffix="b">2008b</bibRefCitation>
) considered it as &quot; 
<emphasis id="0FAF1435FF98792EFCF3249366298DA8" box="[856,1070,465,490]" italics="true" pageId="41" pageNumber="42">species inquirenda</emphasis>
&quot;. Further examination was not possible because all previously collected material has been lost.
</paragraph>
</subSubSection>
</treatment>
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