On the pseudoterrestris species group of the spider genus Coelotes (Araneae, Amaurobiidae)
Author
Wang, Xin-Ping
Author
Griswold, Charles E.
Author
Ubick, Darrell
text
Zootaxa
2009
2313
1
34
journal article
10.5281/zenodo.191946
7553d49b-d6a8-42dc-9128-410ac6536908
1175-5326
191946
Coelotes pseudoyunnanensis
sp. nov.
Figs 88–108
; 156
Material examined:
Holotype
.
CHINA
: Yunnan:
3, Nujiang Prefecture, Pianma Yakou, pass over Gaoligongshan, at elev.
3200 m
,
N25°58'
,
E98°41'
,
October 11, 1998
, C. Griswold, D. Kavanaugh, C.L. Long (
HNU
).
Paratype
.
CHINA
: Yunnan:
3Ƥ, in same vial as
holotype
(
HNU
); 3Ƥ, Nujiang Prefecture, Pianma Yakou, pass over Gaoligongshan,
3200 m
,
N25°58'
,
E98°41'
,
October 11, 1998
, C. Griswold, D. Kavanaugh, C.-L. Long (
CAS
,
CASENT
9029801); 132Ƥ, Gongshan County, Cikai Township, Dabadi troops place along River banks,
3030-3045 m
,
N27°47'
,
E98°30'
,
October 2–3, 2002
, night,
October 8–10, 2002
, H.M. Yan (
CAS
,
CASENT
9025799); 131Ƥ,Gongshan County, Cikai Township,
16.8 km
W of Gongshan on Dulong Valley Road at Heiwadi,
2054-2200 m
,
N27°47'
,
E98°35'
,
October 10, 2002
, H.M. Yan (
HNU
,
CASENT
9025797); 131Ƥ, Gongshan County, Cikai Township,
41 km
W of Gongshan on Dulong Valley Road at Dabadi,
2984 m
,
N27°47'
,
E98°30'
,
October 1, 2002
, H.M. Yan (
CAS
,
CASENT
9025801).
FIGURES 88–91.
Coelotes pseudoyunnanensis
sp. nov.
, female holotype (HNU) from Nujiang, Pianma Yakou.
88,89
Female epigynum, ventral and dorsal view;
90,91
Male palp, ventral and retrolateral view.
Other material examined.
CHINA
: Yunnan:
13,Gongshan County, Cikai Township,
41 km
W of Gongshan on Dulong Valley Road at Dabadi two sides,
3100 m
,
N27°47'
,
E98°32'
,
September 30, 2002
, H.M. Yan (
CAS
,
CASENT
9025802); 1Ƥ, Gongshan County, along Dulong Valley Road about
38–41 km
,
2800– 2900 m
,
N27°47'05.8"
,
E98°31'41.5"
,
September 23, 2002
, X. Xu & C.M. Yin (
HNU
,
CASENT
9025803); 2Ƥ, Gongshan County, Cikai Township,
41 km
W of Gongshan on Dulong Valley Road at Dabadi,
3000 m
,
N27.79655°
,
E098.50562°
,
September 27 to October 6, 2002
, D.H. Kavanaugh, P.E. Marek, D.Z. Dong, & X.C. Liang (
CAS
,
CASENT
9025804); 1Ƥ, Gongshan County, Cikai Township,
41 km
W of Gongshan on Dulong Valley Road at Dabadi two sides,
3100 m
,
N27°27'
,
E98°37'
,
September 23, 2002
, H. M. Yan (
HNU
,
CASENT
9025805); 2Ƥ, Gongshan County, Cikai Township,
41 km
W of Gongshan on Dulong Valley Road at Dabadi,
3000 m
,
N27.79655°
,
E098.50562°
,
September 27 to October 6, 2002
, D.H. Kavanaugh, P.E. Marek, D.Z. Dong, & X.C. Liang (
HNU
,
CASENT
9025806); 1Ƥ, Gongshan County, Cikai Township, Dabadi north bank,
3090 m
,
N27°47'
,
E98°30'
,
September 29, 2002
, H.M. Yan (
CAS
,
CASENT
9025807); 13, Gongshan County, along Dulong Valley Road about
38-41 km
,
2800–2900 m
,
N27°47'05.8"
,
E98°31'41.5"
,
September 23, 2002
, X. Xu & C.M. Yin (
CAS
,
CASENT
9025800); 131Ƥ, Lushui County, Pianma Yakou,
58.1 km
W of Nu Jiang Road on Pianma Road,
3140 m
,
N25.97288°
,
E098.68336°
,
October 15, 2002
, D. H. Kavanaugh, P.E. Marek, & H.B. Liang (
CAS
,
CASENT
9025854); 2Ƥ, Tengchong County, Mingguang Township, Zizhi Village, up slope,
N25.48351°
,
E098.37149°
,
2880 m
; down slope,
N25.47531°
,
E098.37266°
,
2756 m
,
May 19, 2006
, C.M. Yin, J.F. Hu & X.H. Yang (
HNU
,
CASENT
9025624); 13, Tengchong County, Jietou Township,
0.4 km
N of Dahetou, Lingganjiao at Longtang He,
N25.74277°
,
E98.69691°
,
2020 m
,
May 15–20, 2006
, pitfall traps, D. Kavanaugh (
HNU
,
CASENT
9025590); 1Ƥ, Mingguang Township, Zizhi Village,
N25.80894°
,
E098.62080°
,
2890 m
,
May 23, 2006
, X.J. Peng, X.P. Wang & P. Hu (
CAS
,
CASENT
9025592, same vial with 132Ƥ
Coelotes incertus
); 1Ƥ, 00-QF-17 (
CAS
,
CASENT
9025603); 13, Lushui County, Pianma Township, Feng Xue Yakou, elev.
3142 m
,
N26.00949°
,
E98.61704°
, along the road,
May 17, 2005
, K. J. Guo (
HNU
,
CASENT
9022074); 2Ƥ, Longyang County, Bawan District, Nankang Yakou, elev.
2180 m
,
N24.83178°
,
E98.76472°
, understory of good forest on Efacing slope, night collecting,
May 22, 2005
, C. Griswold & D. Kavanaugh (
CAS
,
CASENT
9022287); 1Ƥ, Lushui County, Feng Xue Yakou, E. side on Pianma Road, elev.
3150 m
,
N25.97288°
,
E98.68336°
, hand collecting under rocks,
May 17, 2005
, C. Griswold (
CAS
,
CASENT
9022355,
DNA
sample); 2Ƥ, Gongshan County, Cikai Township,Dabadi troops place south bank of River,
3030 m
,
N27°47'
,
E98°31'
,
September 28, 2002
, H. M. Yan (
CAS
,
CASENT
9025798, same vial with 13
Gnaphosidae
); 1Ƥ, Mingguang Township Zizhi Village,
N25.80894°
,
E098.62080°
,
2890 m
,
May 23, 2006
, road cut, about
2 miles
down from Yakou border station, X.J. Peng, X.P. Wang & P. Hu (
CAS
, in same vial with 2Ƥ13
Draconarius insertus
,
CASENT
9025592); 1Ƥ, Zhongdian County,
51 km
SSE Zhongdian,
2970 m
,
N27°25.3'
,
E99°56.5'
, creek valley, mixed coniferous forest with shrubby vegetation, bamboo, vinegar trap,
August 16–19, 2003
, D.W. Wrase (
SMF
); 1Ƥ, Lijiang Naxi county, E Yulongxue Shan,
30 km
N Lijiang,
2800-2900 m
,
N27°09.0'
,
E100°14.9'
, creek valley, secondary mixed forest,
August 13, 2003
, D.W. Wrase (
SMF
).
FIGURES 92–96.
Coelotes pseudoyunnanensis
sp. nov.
, male and female paratypes (CAS, CASENT9025799) from Gongshan County, Cikai, Dabadi.
92,93
Female epigynum, ventral and dorsal view;
94–96
Male palp, prolateral, ventral and retrolateral view.
Etymology:
The specific name refers to its similarity to
C. yunnanensis
.
Diagnosis:
The female resembles
C. yunnanensis
in having a similar vulva but can be distinguished by the anteriorly originating epigynal teeth and small, laterally extending copulatory ducts (
Figs 88–89
). The male can be recognized by the broad conductor (
Figs 90–91
).
Description:
Male (
holotype
): Medium sized coelotine, total length 6.70 (
Fig. 97
). Carapace 3.50 long, 3.20 wide. Abdomen 3.20 long, 2.00 wide. AME smallest, ¾ the size of other eyes which are subequal or with ALE slightly larger (AME 0.16, ALE 0.20, PME 0.18, PLE 0.19); AME separated by half of their diameter, from ALE by 1/3 of AME diameter; PME separated by slightly more than half of their diameter, from PLE by slightly more than PME diameter (AME-AME 0.07, AME-ALE 0.05, PME-PME 0.10, PME-PLE 0.18, AME-PME 0.13) (
Fig. 98
). Promargin of chelicera with 3 teeth, retromargin 3. Labium slightly wider than long (L/W=0.96) (
Fig. 99
). Palpal patellar apophysis short, broad, distinctly extending and pointing distally; RTA more than half of tibial length, distinctly extending distally; lateral tibial apophysis broad, distinctly separated from RTA; cymbial furrow short, less than half of cymbial length; conductor short, broad, somewhat twisted, with a dorsal apophysis, a small basal lamella; median apophysis broad, round, spoonshaped; embolus short, filiform, originating prolaterally (
Figs 90–91
,
94–96
).
FIGURES 97–102.
Coelotes pseudoyunnanensis
sp. nov.
, male (97–99) and female (100–102) paratype (CAS, CASENT9025799) from Gongshan County, Cikai, Dabadi.
97,100
Habitus, dorsal view;
98,101
Eyes, view between from and dorsal;
99,102
Labia, ventral view.
FIGURES 103–108.
Coelotes pseudoyunnanensis
sp. nov.
, female epigynum, ventral and dorsal view.
103,104
From Tengchong County, Mingguang, Zizhi (HNU, CASENT9025624);
105,106
No detailed label (CAS, CASENT9025603);
107,108
From Gongshan County, Cikai (HNU, CASENT9025797).
Female (one of the
paratypes
measured): Medium sized coelotine, total length 7.70 (
Fig. 100
). Carapace 3.50 long, 2.20 wide. Abdomen 4.20 long, 2.80 wide. AME smallest, ¾ size of lateral eyes which are subequal; PME slightly larger than AME (AME 0.13, ALE 0.17, PME 0.15, PLE 0.17); anterior eyes equally separated by half of AME diameter; PME separated from each other by 2/3 of their diameter, from PLE by slightly more than PME diameter (AME-AME 0.06, AME-ALE 0.05, PME-PME 0.10, PME-PLE 0.17, AME-PME 0.16) (
Fig. 101
). Promargin of chelicera with 3 teeth, retromargin 3. Labium slightly longer than wide (L/W=1.05) (
Fig. 102
). Epigynal teeth short, arise anteriorly at atrium, separated by atrial width; atrium small, with distinct, continuous anterior margin but indistinct lateral margins; copulatory ducts small, slightly extending laterally; spermathecae broad, slightly separated, slightly extending anteriorly; spermathecal heads arising anteriorly from subdistal spermathecae (
Figs 88–89
,
92–93
,
103–108
).
Distribution:
China
(Yunnan) (
Fig. 156
).