<document id="55E35C9AAB3CE59DB08B7B89D4265979" ID-CLB-Dataset="28664" ID-DOI="http://dx.doi.org/10.3897/zookeys.812.30937" ID-GBIF-Dataset="a1496148-c2fe-4f8d-b78f-c31a5fb5ef81" ID-PMC="PMC6328528" ID-Pensoft-Pub="1313-2970-812-93" ID-PubMed="30636912" ID-ZooBank="F105E9A6A4F842209E1798923FC6535F" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1313-2970-812-93" ModsDocOrigin="ZooKeys 812" ModsDocTitle="The Carabidae (Coleoptera) of Shada Al-A’Ala Nature Reserve, Southwestern Saudi Arabia, with description of a new species of Paussinae" checkinTime="1546610372131" checkinUser="pensoft" docAuthor="Abdel-Dayem, Mahmoud S., Elgharbawy, Ali A., Rasool, Iftekhar, Nagel, Peter &amp; Aldhafer, Hathal M." docDate="2019" docId="10A01FFE6D6E62E10A816862A9B37908" docLanguage="en" docName="ZooKeys 812: 93-131" docOrigin="ZooKeys 812" docSource="http://dx.doi.org/10.3897/zookeys.812.30937" docTitle="Harpalus impressus Roth 1851" docType="treatment" docVersion="7" lastPageNumber="103" masterDocId="EA4D5B10923B0773C624D339FFBD422E" masterDocTitle="The Carabidae (Coleoptera) of Shada Al-A'Ala Nature Reserve, Southwestern Saudi Arabia, with description of a new species of Paussinae" masterLastPageNumber="131" masterPageNumber="93" pageNumber="103" updateTime="1701384611363" updateUser="ExternalLinkService">
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<mods:title id="F7EED9E8129DE96C23F9D66CC1EB5C7F">The Carabidae (Coleoptera) of Shada Al-A'Ala Nature Reserve, Southwestern Saudi Arabia, with description of a new species of Paussinae</mods:title>
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<mods:namePart id="9AEE317854331CA65C743EB69609E9BD">Abdel-Dayem, Mahmoud S.</mods:namePart>
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<mods:namePart id="E9C1583ACA9420982F8CBE4EEB3878D0">Elgharbawy, Ali A.</mods:namePart>
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<mods:namePart id="D43EF8060C50572FC400AAB6A6CD5068">Rasool, Iftekhar</mods:namePart>
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<mods:namePart id="9870C5A143A454AA283192A59355D6E0">Nagel, Peter</mods:namePart>
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<mods:namePart id="D18349C6C3BEB8B01E3A4DB4C7FAA2FC">Aldhafer, Hathal M.</mods:namePart>
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<paragraph id="17FDBC6385ACA9A69E4D07E7B7E15C93" pageId="10" pageNumber="103">
<taxonomicName id="6F609DE5C3DD5F3AB7CFC8F385AD4CD3" ID-CoL="3JSLN" authority="Roth, 1851" authorityName="Roth" authorityYear="1851" class="Insecta" family="Carabidae" genus="Harpalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Harpalus impressus" order="Coleoptera" pageId="10" pageNumber="103" phylum="Arthropoda" rank="species" species="impressus">Harpalus impressus Roth, 1851</taxonomicName>
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<subSubSection id="A2C9B9BF5B6ED70BF2BA122540B31D4B" pageId="10" pageNumber="103" type="materials_examined">
<paragraph id="D5FCF55E0679E8487A042B0D2D90F93A" pageId="10" pageNumber="103">Material examined.</paragraph>
<paragraph id="B4FD346861AA8BF5F24C63AA7F494812" pageId="10" pageNumber="103">892 m: 23.IV.2014, LT, 1 ex. 1,474 m: 2.XI.2013, HP, 3 exs.</paragraph>
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<subSubSection id="3AF490613FAF6D2956ADA82651B9A0D3" pageId="10" pageNumber="103" type="distribution">
<paragraph id="18A5FB87E0BA296589378F44CCC90C9E" pageId="10" pageNumber="103">General distribution and zoogeography.</paragraph>
<paragraph id="C40EEA2022127DAA1082CFEAAF19423A" pageId="10" pageNumber="103">ER, ET, SD, SA, YE, UG. AFR species.</paragraph>
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<subSubSection id="F02A1A7FF40B3A9F9DE469EA07810515" pageId="10" pageNumber="103" type="published records">
<paragraph id="BDDCB4D8775AC8BACD4BDB169E8190D5" pageId="10" pageNumber="103">Published records.</paragraph>
<paragraph id="36F93BCC62D7456DCFBAF994D08D3A5D" pageId="10" pageNumber="103">
Asir (Basilewsky 1979; 
<bibRefCitation id="E47606C6929C8251F16C4C76DA40AF66" author="Abdel-Dayem, MS" journalOrPublisher="Zootaxa" pageId="30" pageNumber="123" pagination="341 - 371" title="Faunistic inventory and zoogeographical analysis of the ground beetles (Coleoptera, Carabidae) of Garf Raydah Nature Reserve, Southwestern of Saudi Arabia, and description of a new species of Paussinae." url="https://doi.org/10.11646/zootaxa.4514.3.3" volume="4514" year="2018">Abdel-Dayem et al. 2018</bibRefCitation>
).
</paragraph>
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<subSubSection id="90C13F11E1896F5D2C9AFB95D7777D8F" pageId="10" pageNumber="103" type="remarks">
<paragraph id="873698BBAE2FB91B39E5AFB3F645C9F9" pageId="10" pageNumber="103">Remarks.</paragraph>
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A rare species that was collected during summer and autumn from 
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thorn woodlands. David Wrase identified this species.
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