Revision of Trassedia (Hymenoptera: Ceraphronidae), an Evolutionary Relict With an Unusual Distribution Author Mikó, István Frost Entomological Museum, Department of Entomology, The Pennsylvania State University, University Park, PA, istvan.miko@gmail.com Author Trietsch, Carolyn Frost Entomological Museum, Department of Entomology, The Pennsylvania State University, University Park, PA, Author Kamp, Thomas van de Laboratory for Applications of Synchrotron Radiation, Karlsruhe Institute of Technology (KIT), Kaiserstr. 12, 76131 Karlsruhe, Germany, Author Masner, Lubomír Canadian National Collection of Insects, Arachnids and Nematodes, Central Experimental Farm, Ottawa, ON, Canada, Author Ulmer, Jonah M. Frost Entomological Museum, Department of Entomology, The Pennsylvania State University, University Park, PA, Author Yoder, Matthew Jon Illinois Natural History Survey, University of Illinois at Urbana-Champaign, Champaign, IL, Author Zuber, Marcus Laboratory for Applications of Synchrotron Radiation, Karlsruhe Institute of Technology (KIT), Kaiserstr. 12, 76131 Karlsruhe, Germany, Author Sandall, Emily L. Frost Entomological Museum, Department of Entomology, The Pennsylvania State University, University Park, PA, Author Baumbach, Tilo Laboratory for Applications of Synchrotron Radiation, Karlsruhe Institute of Technology (KIT), Kaiserstr. 12, 76131 Karlsruhe, Germany, Author Deans, Andrew R. Frost Entomological Museum, Department of Entomology, The Pennsylvania State University, University Park, PA, text Insect Systematics and Diversity 2018 AIFB 2018-12-04 2 6 4 1 29 http://dx.doi.org/10.1093/isd/ixy015 journal article 10.1093/isd/ixy015 2399-3421 7168382 Trassedia yanegai Mikó and Trietsch sp. nov. (Zoobank LSID: urn:lsid:zoobank. org:act: 53D5E575-4ED3-4146-A7F2-19ECF032846E ) Diagnosis Trassedia yanegai shares the short setae on the dorsal region of the cranium (shorter than the diameter of the median ocellus; Fig. 3C and D ), the reduced anterior ocellar fovea that does not extend to the antennal scrobe ( Fig. 3C and D ), the punctate ocular impression along the inner orbit (io: Fig. 3C and D ) and a wide Waterston’s evaporatorium that reaches the lateral 13 of the tergite (wop: Fig. 4A and B ) with T. luapi and T. australiensis . Trassedia yanegai differs from these species in the lack of the notaulus, the lack of scutes on the vertex, the anteriorly gradually widening preoccipital sulcus (pof: Fig. 3C ), the foveolate sternaulus (ster: Fig. 5C ) and the large eyes (HW/IOS = 3.7, Fig. 3C ). The female pedicel and F1 are brown and F2–F4 are whitish in T. yanegai , while the female pedicel and F1–F3 are yellow and F4–F9 are dark brown in T. luapi . Description Body length: 3.95 mm . Color hue pattern female: cranium, mesosomablack, F5–F9 black, F2–F4 white, scape, legs, pedicel, F1 ochre. Color intensity pattern female: pedicel, F1, fore leg, mesocoxa darker than rest of middle leg, hind leg, scape. Structure of scutes on head and mesosoma: scute surface on head and mesosoma flat, scutes indistinct. Head: HW:HH = 1.1. HW/IOS Female: 3.8. Maximum eye diameter versus minimum eye diameter: 1.2. Interommatidial seta length: interommatidial seta length less than facet diameter. Occipital carina medially: continuous medially. Seta length on dorsal region of cranium versus diameter of median ocellus: shorter. Scutes on vertex count: absent. Preoccipital furrow anterior extension: adjacent anteriorly to the posterior margin of the median ocellus. Preoccipital furrow anterior region versus posterior region sculpture: posterior region crenulate, anterior region smooth or finely reticulate. Preoccipital furrow anterior region width versus posterior region width: wider anteriorly than posteriorly. Female OOL: POL: LOL: 0.1:0.5:1.0. Preocellar pit count: absent. Carina delimiting antennal scrobe count: absent. Transverse striation on upper face count: present. Anterior ocellar fovea shape: fovea not extended ventrally into facial sulcus. Supraclypeal depression count: present. Ocular impression sculpture: punctate (fovea of ocellar impression are well separated from each other). Antenna: Female scape length versus pedicel length: 2.4. Female F1 length versus pedicel length: 1.1. Female ninth flagellomere length: F9 less than F7+F8. Mesosoma: Mesosoma shape: not compressed laterally, as wide as high or wider than high. Pronope count: absent. Anterior slope of mesonotum shape: Anterior slope of mesonotum at obtuse angle to dorsal surface of mesonotum in lateral view. Antero-admedian line count: present. Notaulus count: absent. Notaulus anterior origin versus anterolateral angle of mesoscutum (ball-and-socket articulation between pronotum and mesoscutum): Notaulus arises medially of anterolateral angle of the mesoscutum. Scutes on posterior region of mesoscutum and dorsal region of mesoscutellum count: present. Epicnemial carina count: complete. Sternaulus count: present. Sternaulus sculpture: scalloped. Anterior metapleural carina count: present. Carina limiting posteriorly antecosta count: present. Lateral propodeal carina count: present. Wings: Stigmal vein length versus pterostigma marginal length: stigmal vein longer than the pterostigma marginal length. Etymology The species epithet refers to Douglas Yanega (Senior Museum Scientist of the Entomology Research Museum, University of California , Riverside), who drew our attention to the holotype of the new species.