Order Chiroptera - Family Phyllostomidae Author Wilson, Don E. Author Reeder, DeeAnn text 2005 The Johns Hopkins University Press Baltimore Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 1 395 426 book chapter 0-8018-8221-4 10.5281/zenodo.7316519 Lophostoma d'Orbigny 1836 Lophostoma d'Orbigny 1836 , Voy. Am. Merid., Atlas Zool., Vol. 4: 11 . Type Species: Lophostoma silvicolum d'Orbigny 1836 Synonyms: . Species and subspecies: 5 species with 4 subspecies: Species Lophostoma brasiliense Peters 1866 Species Lophostoma carrikeri J. A. Allen 1910 Species Lophostoma evotis Davis and Carter 1978 Species Lophostoma schulzi Genoways and Williams 1980 Species Lophostoma silvicolum d'Orbigny 1836 Subspecies Lophostoma silvicolum subsp. silvicolum d'Orbigny 1836 Subspecies Lophostoma silvicolum subsp. centralis Davis and Carter 1978 Subspecies Lophostoma silvicolum subsp. laephotis Thomas 1910 Subspecies Lophostoma silvicolum subsp. occidentalis Davis and Carter 1978 Discussion: Formerly included Tonatia , but see Lee et al. (2002) , who demonstrated that Tonatia as traditionally defined is not monophyletic. Those authors proposed restricting Tonatia to the type species and its close relative ( bidens and saurophila ), and using the next available generic name ( Lophostoma ) for the remaining species, which together form a clade that is not closely related to Tonatia . That recommendation is followed here. Keys to species now included in Lophostoma were provided by Genoways and Williams (1984) and Medellín and Arita (1989).