Food Relocation Behavior and Synopsis of the Southern South American Genus Glyphoderus Westwood (Scarabaeidae: Scarabaeinae: Eucraniini) Author Ocampo, Federico C. text The Coleopterists Bulletin 2004 2004-06-30 58 2 295 305 http://dx.doi.org/10.1649/685 journal article 10.1649/685 1938-4394 10104682 Glyphoderus sterquilinus ( Westwood, 1837 ) ( Figs. 5 , 6 ) Glyphoderus sterquilinus ( Westwood, 1837 ) :13. Anomiopsis sterquilinus Westwood, 1837:13 . Original combination. Diagnosis. Males. This species is separated from other Glyphoderus species by the shape and placement of the pronotal horns and tubercles ( Fig. 5 ). Glyphoderus sterquilinus has the pronotum with two horns in middle and two transversally elongated tubercles (sometimes developed as horns) on basal half; horns on margin of central depression; tubercles of basal half included in central depression; apical medial horn with ‘‘arms’’ robust, with anteromedial notch. Some specimens of G. sterquilinus have the posterior pronotal tubercles on the central depression margin (observed particularly in minor males), in this case, the shape of the medial horn will separate this species from G. monticola ( Figs. 4 , 5 ). No morphological characters were found to diagnose females of this species, females can be identified only by association with males. Distribution ( Fig. 6 ). ARGENTINA : Mendoza : Las Catitias ( 7 km S) (6); Santa Rosa (1); Villavicencio (1). La Rioja : Chepes (2); Ulapes ( 17 km S) (28); Ulapes ( 10 km N) (8). San Juan : Encón ( 12 km E) (1); Nikizanga (2); Morayes –Astica road (1). San Luis : Jarilla (1); La Chañarienta (2); San Antonio (5); Talita (8); no more data (11) . Santiago del Estero : No more data (6).