Revision of the Hylicellidae of the Late Triassic of Queensland (Hemiptera Cicadomorpha: Hylicelloidea)
Author
Lambkin, Kevin J.
text
Zootaxa
2020
2020-06-12
4790
3
525
539
journal article
10.11646/zootaxa.4790.3.7
1175-5326
3891303
F011C13E-8E90-45D8-ABA3-B7D58582648A
Mesocixiodes
Tillyard, 1922
Mesocixiodes
Tillyard, 1922: 462
.
Type
species.
Mesocixiodes termioneura
Tillyard, 1922
, by original designation.
Emended diagnosis.
Tegmen
c
. 3.5 times longer than wide, finely to somewhat coarsely punctate and tending to irregularly transversely rugose, extreme apical area possibly less so; costal margin broadly rounded, apex narrowly rounded; peripheral membrane of even width; costal space
c
. 3 times as wide as medial cell; R and M separating near apex of basal cell; basal cell quite broad proximally, slightly tapering and closed distally by a fusion for a short length of M and CuA; primary fork of R just before tegmen length, far basal to primary fork of M which is well beyond primary fork of RA and just proximal to very shallow CuA fork; RA
1
simple, obliquely angled towards anterior margin; RA
2
stem directed apically, more or less parallel with costal margin, with 4 or 5 long, sub-equal pectinate branches directed antero-apically; RA
1
+ RA
2
field occupying entire antero-apical margin; RP deeply forked; one
ir
, running from RA
2
to RP
1
; RP
2
fused with M
1+2
for a short length (
RK1
and
RK2
), or separate, with a single
r-m
, in line with
ir
(
holotype
); M with 4 or 5 branches;
im
between M
2
and M
3
or M
3+4
;intra-medial cell quite broad but very small,
c
. 2 times longer than wide, occupying a very small portion of post-nodal area; CuA branches short, CuA
1
strongly angled at
m-cua
which joins M
3+4
around midpoint of intra-medial cell; crossvein field tending to alignment at level of claval apex, thus creating a fairly even series of long, sub-equal apical cells; clavus not preserved
.
Notes.
Mesocixiodes
was placed by
Tillyard (1922
,
1926
),
Becker-Migdisova (1962a)
, and
Jell (2004)
in
Cixiidae (Fulgoroidea)
, by
Handlirsch (1939)
in
Fulgoridae
, by
Evans (1956)
in
Chiliocyclidae
, and was considered by
Carpenter (1992)
as ‘Family uncertain’. It was transferred to
Hylicellidae
by
Hamilton (1992)
. It differs from all other hylicellids in the very late forking of M, well distal of that of RA, the very small intra-medial cell, and the tendency to alignment of the crossvein field at the claval apex to create a fairly even series of long, sub-equal apical cells.