Growth, inter- and intraspecific variation, palaeobiogeography, taphonomy and systematics of the Cenozoic ghost shrimp Glypturus Author Klompmaker, A. Florida Museum of Natural History, University of Florida, 1659 Museum Road, PO Box 117800, Gainesville, FL 32611, USA; Author Hy ̆ z, Matu ̆ s Author , Roger W. Port Author Kowalew, nd Michał text Journal of Systematic Palaeontology 2016 2015-03-16 14 2 99 126 http://dx.doi.org/10.1080/14772019.2015.1009505 journal article 10.1080/14772019.2015.1009505 1478-0941 PMC4724948 26819570 C7F0C071-F2AD-4684-B277-037B6F91BF0E Glypturus toulai ( Rathbun, 1919 ) ( Fig. 16 ) ‡ 1919 Callianassa toulai Rathbun , 146. 2013 Glypturus toulai (Rathbun) ; Hy̆zńy et al .: 132, figs 4, 5 (cum syn.). Diagnosis. Moderately tuberculated Glypturus ; lateral tuberculation on outer side of propodus extending from near base of fixed finger diagonally to lower margin and reaching proximal lower corner; lateral tuberculation on inner side of the propodus restricted mainly to area of articulation with dactylus in small specimens, but covering [central portion of inner side of manus in large specimens.] (modified from Hy̆zńy et al. 2013, pp. 132 — 133). Material. UF 203348 ( Fig. 16E, F ). NHMW 1933/0018/ 0159 ( Fig. 16A — D ), 1933/0018/0160 ( Fig. 16G — J + 3 fixed fingers). BMNPH PI IC 395 ( Fig. 16K — M ). Occurrence. This species is known from the upper Miocene Gatun Formation in Panama (Supplemental Table 1 ). Description. Manus length exceeding height (length/ height ratio ~1.1). Upper margin converging distally, proximally keeled, bearing three prominent spines pointing distally, proximalmost spine around mid-margin, keel terminating in blunt corner. Lower margin sharp and keeled, in larger specimens denticulate, lined with a row of setal pits on inner lateral side. Proximal margin convex on outer face, concave on inner, [heightwise groove near proximal margin concave on inner side, concave to straight on outer side]. Distal margin weakly convex. Lateral tuberculation on outer side from base of fixed finger diagonally to lower margin and proximal lower corner. Lateral tuberculation on inner side of propodus restricted mainly to area of articulation with dactylus in small specimens, but covering [central portion of inner side of manus in large specimens]. Fixed finger triangular, sharply pointed, with distinct blunt tooth on occlusal margin around mid-length, tooth pointing distally. Dactylus stout, occlusal margin [with or] without tooth [around mid-length], tip hooked (modified from Hy̆zńy et al. 2013, pp. 134 — 135).