A revision of the genus Cheliplana de Beauchamp, 1927 (Rhabdocoela: Schizorhynchia), with the description of six new species
Author
Gobert, Stefan
0000-0003-1949-9174
Hasselt University, Centre for Environmental Sciences, Research Group Zoology: Biodiversity and Toxicology, Universitaire Campus Gebouw D, B- 3590 Diepenbeek, Belgium. & stefan. gobert @ uhasselt. be; https: // orcid. org / 0000 - 0003 - 1949 - 9174
stefan.gobert@uhasselt.be
Author
Diez, Yander L.
0000-0001-8741-4799
Hasselt University, Centre for Environmental Sciences, Research Group Zoology: Biodiversity and Toxicology, Universitaire Campus Gebouw D, B- 3590 Diepenbeek, Belgium. & University of Oriente, Departement of Biology and Geography, Avenida Patricio Lumumba s / n, CP 90500, Santiago de Cuba, Cuba. Beaty Biodiversity Research Centre, Departments of Botany and Zoology, University of British Columbia, 3529 - 6270 University Blvd, Vancouver, BC, V 6 T 1 Z 4 Canada. & yander @ uo. edu. cu; https: // orcid. org / 0000 - 0001 - 8741 - 4799
yander@uo.edu.cu
Author
Monnens, Marlies
0000-0002-9133-1512
Hasselt University, Centre for Environmental Sciences, Research Group Zoology: Biodiversity and Toxicology, Universitaire Campus Gebouw D, B- 3590 Diepenbeek, Belgium. & marlies. monnens @ uhasselt. be; https: // orcid. org / 0000 - 0002 - 9133 - 1512
marlies.monnens@uhasselt.be
Author
Reygel, Patrick
0000-0001-8721-6290
Hasselt University, Centre for Environmental Sciences, Research Group Zoology: Biodiversity and Toxicology, Universitaire Campus Gebouw D, B- 3590 Diepenbeek, Belgium. & patrick. reygel @ uhasselt. be; https: // orcid. org / 0000 - 0001 - 8721 - 6290
patrick.reygel@uhasselt.be
Author
Van Steenkiste, Niels W. L.
0000-0002-2676-7862
niels. vansteenkiste @ botany. ubc. ca; https: // orcid. org / 0000 - 0002 - 2676 - 7862
niels.vansteenkiste@botany.ubc.ca
Author
Leander, Brian S.
0000-0003-0798-0470
bleander @ mail. ubc. ca; https: // orcid. org / 0000 - 0003 - 0798 - 0470
bleander@mail.ubc.ca
Author
Artois, Tom
Hasselt University, Centre for Environmental Sciences, Research Group Zoology: Biodiversity and Toxicology, Universitaire Campus Gebouw D, B- 3590 Diepenbeek, Belgium.
text
Zootaxa
2021
2021-05-17
4970
3
453
494
journal article
6258
10.11646/zootaxa.4970.3.2
281c9017-f55b-47bf-bea0-e511a7173605
1175-5326
4766692
FEABE248-E1EA-48F5-A1AF-0077FE40C257
Cheliplana setosa
Evdonin, 1971
Fig. 9A–D
Material examined. New material.
CANADA
• 1 whole mount;
British Columbia
,
Calvert Island
,
Foggy Cove
;
51°39’07”N
,
128°08’32”W
;
9 Apr. 2016
; medium-grained sand in between boulders in the lower intertidal; MI4193
• 1 whole mount;
British Columbia
, Bamfield, Brady Beach;
48°49’40”N
,
125°09’12”W
;
3 Jun. 2015
; mediumgrained sand in lower intertidal; MI4194
.
Reference material.
1 whole-mounted specimen (
SMNH
90403) and 1 serially sectioned specimen (
SMNH
102990) from
California
,
United States
.
FIGURE 9
.
A–D.
Cheliplana setosa
Evdonin, 1971
from British Columbia, Canada.
A.
Live specimen.
B.
Detail of the male copulatory organs in a live specimen.
C.
Detail of the proboscis with proboscis hooks and hook supports in a wholemounted specimen.
D.
Detail of the male copulatory bulb with prostate vesicle and tripartite cirrus in a whole-mounted specimen.
E.
Cheliplana californica
Karling, 1989
from British Columbia, Canada. Detail of the male copulatory bulb with cirrus in the whole-mounted specimen. Scale bars: A = 100 μm; B = 20 μm; C = 10 μm; D–E = 20 μm.
Known distribution
. Posyet, Peter the Great Gulf,
Russia
(
Evdonin 1977
). Monterey Bay,
California
,
United States
(
Karling 1983
). Ría de Villaviciosa, near San Martin,
Spain
(
Noreña
et al.
2007
).
Remarks
. Live specimens are ~
1.5 mm
long. Specimens are yellowish to pinkish in colour (
Fig. 9A
). The proboscis bears curved proboscis hooks measuring 18–20 μm (h,
Fig. 9C
). Hook supports are about 12 μm long in the specimens from
Canada
(hs,
Fig. 9C
). Rod-shaped, weakly sclerotised proboscis sidepieces were observed by
Karling (1983)
. The prepharyngeal tube is armed with spines (ppc,
Fig. 9A
).
A single, large testis is situated alongside the pharynx (ph, t,
Fig. 9A
).
Karling (1983)
reports a lobe containing sperm at the posterior end of the testis. A pair of seminal vesicles, one of which ending blindly, the other connecting to the testis via an unpaired vas deferens, enter the proximal end of the copulatory bulb (cb, vs,
Fig. 9A–B,D
). The copulatory bulb is oviform (cb,
Fig. 9A–B,D
). In the Russian specimens,
Evdonin (1971)
measures a size range of 110 to 120 μm for the copulatory bulb. The Californian specimen described by
Karling (1983)
possesses a copulatory bulb of ~130 μm. The copulatory bulbs in the new specimens from
Canada
are somewhat larger still, measuring 130–160 μm. The copulatory bulb holds a cirrus of 40–60 μm (
Russia
and California) (
Evdonin 1971
;
Karling 1983
) or 68–85 μm (
Canada
) (ci,
Fig. 9 A–B,D
). The cirrus can be divided in three clearly defined regions: (1) a proximal tube-like region of ~48 μm (California) to 48–60 μm (
Canada
), armed with fine spines, (2) a ring of five or six large spines of about 10–15 μm (
Canada
) long, set in sclerotised pockets, and (3) a distal region of ~7 μm (California) to 15–17 μm (
Canada
) long, wider than the proximal region, armed with hair-like spines.
A single ovary (ov,
Fig. 9A
) and bursa are situated near the caudal body end.