<document ID-DOI="http://doi.org/10.5281/zenodo.3792844" ID-GBIF-Dataset="142a25ed-47c3-4361-bc24-6f1b3e8dba5c" ID-GBIF-Taxon="163755041" ID-ISSN="1313–2970" ID-Zenodo-Dep="3792844" ID-ZooBank="664C49F1-5384-43C4-8BF1-CE76AC11D32E" approvalRequired="10" approvalRequired_for_treatments="10" checkinTime="1587998787238" checkinUser="plazi" docAuthor="Klimaszewski, Jan, Savard, Karine, Pelletier, Georges &amp; Webster, Reginald" docDate="2008" docId="D43E034BFFA84159FF17FCE86501FB51" docLanguage="en" docName="ZK_article_1943.pdf" docOrigin="ZooKeys 2 (2)" docStyle="DocumentStyle:BEF2356135E79670CA200BCAD40C4E3A:zookeys.2008.journal_article" docStyleId="BEF2356135E79670CA200BCAD40C4E3A" docStyleName="zookeys.2008.journal_article" docTitle="Gnypeta nigrella Thomson" docType="treatment" docVersion="8" lastPageNumber="64" masterDocId="28077B33FF9D416CFF9BFFC76718FFEC" masterDocTitle="Species review of the genus Gnypeta Thomson from Canada, Alaska and Greenland (Coleoptera, Staphylinidae, Aleocharinae): systematics, bionomics and distribution" masterLastPageNumber="84" masterPageNumber="11" pageId="53" pageNumber="64" updateTime="1643574247635" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:title>Species review of the genus Gnypeta Thomson from Canada, Alaska and Greenland (Coleoptera, Staphylinidae, Aleocharinae): systematics, bionomics and distribution</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Klimaszewski, Jan</mods:namePart>
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
<mods:affiliation>Natural Resources Canada, Canadian Forest Service, Laurentian Forestry Centre, Quebec, QC, Canada &amp; C 136 Eddc &amp; A- 86 C</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Savard, Karine</mods:namePart>
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
<mods:affiliation>Natural Resources Canada, Canadian Forest Service, Laurentian Forestry Centre, Quebec, QC, Canada &amp; C 136 Eddc &amp; A- 86 C</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Pelletier, Georges</mods:namePart>
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
<mods:affiliation>Natural Resources Canada, Canadian Forest Service, Laurentian Forestry Centre, Quebec, QC, Canada &amp; C 136 Eddc &amp; A- 86 C</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Webster, Reginald</mods:namePart>
<mods:affiliation>24 Millstream Drive, Fredericton, NB, Canada &amp; F-</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2008</mods:date>
<mods:detail type="pubDate">
<mods:number>2008-09-04</mods:number>
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<mods:detail type="volume">
<mods:number>2</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
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<mods:start>11</mods:start>
<mods:end>84</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3897/zookeys.2.4</mods:identifier>
<mods:identifier type="GBIF-Dataset">142a25ed-47c3-4361-bc24-6f1b3e8dba5c</mods:identifier>
<mods:identifier type="ISSN">1313–2970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576410</mods:identifier>
<mods:identifier type="ZooBank">664C49F1-5384-43C4-8BF1-CE76AC11D32E</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3792844" ID-GBIF-Taxon="163755041" ID-Zenodo-Dep="3792844" LSID="urn:lsid:plazi:treatment:D43E034BFFA84159FF17FCE86501FB51" httpUri="http://treatment.plazi.org/id/D43E034BFFA84159FF17FCE86501FB51" lastPageNumber="64" pageId="53" pageNumber="64">
<subSubSection box="[140,577,815,842]" pageId="53" pageNumber="64" type="nomenclature">
<paragraph blockId="53.[140,577,815,842]" box="[140,577,815,842]" pageId="53" pageNumber="64">
<heading box="[140,577,815,842]" fontSize="11" level="2" pageId="53" pageNumber="64" reason="3">
<emphasis bold="true" box="[140,577,815,842]" pageId="53" pageNumber="64">
V. 
<emphasis bold="true" box="[170,384,815,842]" italics="true" pageId="53" pageNumber="64">
<taxonomicName authority="Thomson" authorityName="Thomson" box="[170,280,815,842]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="64" phylum="Arthropoda" rank="genus" species="nigrella">Gnypeta</taxonomicName>
nigrella
</emphasis>
species group
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="53" pageNumber="64" type="description">
<paragraph blockId="53.[140,1108,870,1108]" pageId="53" pageNumber="64">
The 
<taxonomicName box="[190,271,870,896]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="64" phylum="Arthropoda" rank="species" species="nigrella">
<emphasis box="[190,271,870,896]" italics="true" pageId="53" pageNumber="64">nigrella</emphasis>
</taxonomicName>
species group may be characterized by the following combination of characters: spermatheca club-shaped, capsule approximately spherical and constricted to a narrow duct connected to a narrow, tubular and sinuate stem (
<figureCitation box="[880,983,941,967]" captionStart="Figs 164-171" captionStartId="54.[140,184,1590,1612]" captionTargetBox="[149,1086,244,1506]" captionTargetId="figure@54.[140,1108,232,1523]" captionTargetPageId="54" captionText="Figs 164-171. Genital structures of Gnypeta nigrella: 164, median lobe of the aedeagus in lateral view, 165, dorsal view; 166, paramere; 167, male tergite 8; 168, male sternite 8; 169, spermatheca; 170, female tergite 8; 171, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774126" httpUri="https://zenodo.org/record/3774126/files/figure.png" pageId="53" pageNumber="64">Figs 169</figureCitation>
, 
<figureCitation box="[996,1042,941,967]" captionStart="Figs 172-180" captionStartId="56.[140,184,1590,1612]" captionTargetBox="[142,1108,243,1519]" captionTargetId="figure@56.[140,1108,242,1532]" captionTargetPageId="56" captionText="Figs 172-180. Genital structures of Gnypeta saccharina: 172, median lobe of the aedeagus in lateral view, 173, dorsal view, 174, apical part of median lobe in ventral view; 175, paramere; 176, male tergite 8; 177, male sternite 8; 178, spermatheca; 179, female tergite 8; 180, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774128" httpUri="https://zenodo.org/record/3774128/files/figure.png" pageId="53" pageNumber="64">178</figureCitation>
, 
<figureCitation captionStart="Figs 181-189" captionStartId="59.[140,184,1590,1612]" captionTargetBox="[152,1108,234,1496]" captionTargetId="figure@59.[140,1109,234,1545]" captionTargetPageId="59" captionText="Figs 181-189. Genital structures of Gnypeta minuta: 181, median lobe of the aedeagus in lateral view, 182, dorsal view; 183, paramere; 184, male tergite 8; 185, male sternite 8; 186, 187, spermatheca; 188, female tergite 8; 189, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774130" httpUri="https://zenodo.org/record/3774130/files/figure.png" pageId="53" pageNumber="64">187, 188</figureCitation>
); median lobe of aedeagus with moderately large bulbus which is streamlined with tubus in dorsal view (
<figureCitation box="[441,542,1011,1038]" captionStart="Figs 164-171" captionStartId="54.[140,184,1590,1612]" captionTargetBox="[149,1086,244,1506]" captionTargetId="figure@54.[140,1108,232,1523]" captionTargetPageId="54" captionText="Figs 164-171. Genital structures of Gnypeta nigrella: 164, median lobe of the aedeagus in lateral view, 165, dorsal view; 166, paramere; 167, male tergite 8; 168, male sternite 8; 169, spermatheca; 170, female tergite 8; 171, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774126" httpUri="https://zenodo.org/record/3774126/files/figure.png" pageId="53" pageNumber="64">Figs 165</figureCitation>
, 
<figureCitation box="[555,601,1011,1037]" captionStart="Figs 172-180" captionStartId="56.[140,184,1590,1612]" captionTargetBox="[142,1108,243,1519]" captionTargetId="figure@56.[140,1108,242,1532]" captionTargetPageId="56" captionText="Figs 172-180. Genital structures of Gnypeta saccharina: 172, median lobe of the aedeagus in lateral view, 173, dorsal view, 174, apical part of median lobe in ventral view; 175, paramere; 176, male tergite 8; 177, male sternite 8; 178, spermatheca; 179, female tergite 8; 180, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774128" httpUri="https://zenodo.org/record/3774128/files/figure.png" pageId="53" pageNumber="64">173</figureCitation>
, 
<figureCitation box="[614,661,1011,1037]" captionStart="Figs 181-189" captionStartId="59.[140,184,1590,1612]" captionTargetBox="[152,1108,234,1496]" captionTargetId="figure@59.[140,1109,234,1545]" captionTargetPageId="59" captionText="Figs 181-189. Genital structures of Gnypeta minuta: 181, median lobe of the aedeagus in lateral view, 182, dorsal view; 183, paramere; 184, male tergite 8; 185, male sternite 8; 186, 187, spermatheca; 188, female tergite 8; 189, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774130" httpUri="https://zenodo.org/record/3774130/files/figure.png" pageId="53" pageNumber="64">182</figureCitation>
), the internal sac with two pronounced subapical sclerites (
<figureCitation box="[354,514,1046,1073]" captionStart="Figs 164-171" captionStartId="54.[140,184,1590,1612]" captionTargetBox="[149,1086,244,1506]" captionTargetId="figure@54.[140,1108,232,1523]" captionTargetPageId="54" captionText="Figs 164-171. Genital structures of Gnypeta nigrella: 164, median lobe of the aedeagus in lateral view, 165, dorsal view; 166, paramere; 167, male tergite 8; 168, male sternite 8; 169, spermatheca; 170, female tergite 8; 171, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774126" httpUri="https://zenodo.org/record/3774126/files/figure.png" pageId="53" pageNumber="64">Figs 164, 165</figureCitation>
, 
<figureCitation box="[527,632,1047,1073]" captionStart="Figs 172-180" captionStartId="56.[140,184,1590,1612]" captionTargetBox="[142,1108,243,1519]" captionTargetId="figure@56.[140,1108,242,1532]" captionTargetPageId="56" captionText="Figs 172-180. Genital structures of Gnypeta saccharina: 172, median lobe of the aedeagus in lateral view, 173, dorsal view, 174, apical part of median lobe in ventral view; 175, paramere; 176, male tergite 8; 177, male sternite 8; 178, spermatheca; 179, female tergite 8; 180, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774128" httpUri="https://zenodo.org/record/3774128/files/figure.png" pageId="53" pageNumber="64">177, 178</figureCitation>
, 
<figureCitation box="[645,751,1047,1073]" captionStart="Figs 181-189" captionStartId="59.[140,184,1590,1612]" captionTargetBox="[152,1108,234,1496]" captionTargetId="figure@59.[140,1109,234,1545]" captionTargetPageId="59" captionText="Figs 181-189. Genital structures of Gnypeta minuta: 181, median lobe of the aedeagus in lateral view, 182, dorsal view; 183, paramere; 184, male tergite 8; 185, male sternite 8; 186, 187, spermatheca; 188, female tergite 8; 189, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774130" httpUri="https://zenodo.org/record/3774130/files/figure.png" pageId="53" pageNumber="64">181, 182</figureCitation>
); female sternite 8 emarginated medially (
<figureCitation box="[251,351,1082,1108]" captionStart="Figs 164-171" captionStartId="54.[140,184,1590,1612]" captionTargetBox="[149,1086,244,1506]" captionTargetId="figure@54.[140,1108,232,1523]" captionTargetPageId="54" captionText="Figs 164-171. Genital structures of Gnypeta nigrella: 164, median lobe of the aedeagus in lateral view, 165, dorsal view; 166, paramere; 167, male tergite 8; 168, male sternite 8; 169, spermatheca; 170, female tergite 8; 171, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774126" httpUri="https://zenodo.org/record/3774126/files/figure.png" pageId="53" pageNumber="64">Figs 171</figureCitation>
, 
<figureCitation box="[364,411,1082,1108]" captionStart="Figs 172-180" captionStartId="56.[140,184,1590,1612]" captionTargetBox="[142,1108,243,1519]" captionTargetId="figure@56.[140,1108,242,1532]" captionTargetPageId="56" captionText="Figs 172-180. Genital structures of Gnypeta saccharina: 172, median lobe of the aedeagus in lateral view, 173, dorsal view, 174, apical part of median lobe in ventral view; 175, paramere; 176, male tergite 8; 177, male sternite 8; 178, spermatheca; 179, female tergite 8; 180, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774128" httpUri="https://zenodo.org/record/3774128/files/figure.png" pageId="53" pageNumber="64">180</figureCitation>
, 
<figureCitation box="[422,470,1082,1108]" captionStart="Figs 181-189" captionStartId="59.[140,184,1590,1612]" captionTargetBox="[152,1108,234,1496]" captionTargetId="figure@59.[140,1109,234,1545]" captionTargetPageId="59" captionText="Figs 181-189. Genital structures of Gnypeta minuta: 181, median lobe of the aedeagus in lateral view, 182, dorsal view; 183, paramere; 184, male tergite 8; 185, male sternite 8; 186, 187, spermatheca; 188, female tergite 8; 189, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774130" httpUri="https://zenodo.org/record/3774130/files/figure.png" pageId="53" pageNumber="64">189</figureCitation>
).
</paragraph>
<paragraph blockId="53.[140,1107,1152,1213]" pageId="53" pageNumber="64">
<emphasis bold="true" box="[140,338,1152,1178]" pageId="53" pageNumber="64">Species included</emphasis>
: 
<taxonomicName authority="Klimaszewski and Webster" authorityName="Klimaszewski and Webster" authorityYear="2008" box="[353,781,1152,1178]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="64" phylum="Arthropoda" rank="species" species="minuta">
<emphasis box="[353,469,1152,1178]" italics="true" pageId="53" pageNumber="64">G. minuta</emphasis>
Klimaszewski and Webster
</taxonomicName>
, 
<taxonomicName authority="LeConte" authorityName="LeConte" box="[793,1020,1152,1178]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="64" phylum="Arthropoda" rank="species" species="nigrella">
<emphasis box="[793,910,1152,1178]" italics="true" pageId="53" pageNumber="64">G. nigrella</emphasis>
LeConte
</taxonomicName>
, 
<taxonomicName authority="Klimaszewski and Webster." authorityName="Klimaszewski and Webster." class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="64" phylum="Arthropoda" rank="species" species="saccharina">
<emphasis italics="true" pageId="53" pageNumber="64">G. saccharina</emphasis>
Klimaszewski and Webster.
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>