The reclassification of Sophiodela and other tiger beetles (Coleoptera, Cicindelidae) based on the structure of the everted internal sac of the male genitalia Author Fukuda, Yuki Kurayoshi, 111 - 8, Kamogawa-town, Tottori, 682 - 0923 Japan E-mail: yuki 03 go 3 @ gmail. com yuki03go3@gmail.com Author Ogawa, Ryo Matsuyama, 6 - 5 - 8, Hakusuidai, Ehime, 791 - 0113 Japan E-mail: ailmasbl 0854 @ yahoo. co. jp Author Hori, Michio Wakayama, 6 - 1 - 2 Takajyomachi, Wakayama, 640 - 8135 Japan E-mail: hori @ terra. zool. kyoto-u. ac. jp text Zootaxa 2019 2019-08-28 4661 2 271 308 journal article 21156 10.11646/zootaxa.4661.2.3 6c12aca1-7350-4265-9ded-6d415152f181 1175-5326 3998528 95722A8A-E02C-47E5-AA9B-777B6CA226E6 Cicindela ( Pancallia ) angulicollis Horn, 1900 ( Figs 5D , 11D ) Cicindela ( Calochroa ) angulicollis Horn, 1900:209 . Cicindela ( Pancallia ) angulicollis : Rivalier 1961:124 . Specimens examined. (N = 1) [ South India ] 1♂ ( MHJ ), Madras State, Coimbatore, X. 1979 , G. Duranto leg. Description. Internal sac everted apically in lateral view, parallel to median lobe. VA largest in bladders; DA and VB the second largest; DLL smallest in bladders. VA roundly swollen ventrally. DA everted dorsally. DLL roundly swollen, vested numerously with spines, which are weakly chitinous. VB oval and widely swollen, everted left laterally, vested numerously with long spines, which are forming a band. Flagellum well short, curved. Medial tooth very short, extending ventro-apically, tapered gently at apex. Remarks . This species shares the characteristics of the position of each bladder with C . ( P ). princeps , but is distinguished by the following characteristics: DLL small; B non-existent; medial tooth very short.