The reclassification of Sophiodela and other tiger beetles (Coleoptera, Cicindelidae) based on the structure of the everted internal sac of the male genitalia Author Fukuda, Yuki Kurayoshi, 111 - 8, Kamogawa-town, Tottori, 682 - 0923 Japan E-mail: yuki 03 go 3 @ gmail. com yuki03go3@gmail.com Author Ogawa, Ryo Matsuyama, 6 - 5 - 8, Hakusuidai, Ehime, 791 - 0113 Japan E-mail: ailmasbl 0854 @ yahoo. co. jp Author Hori, Michio Wakayama, 6 - 1 - 2 Takajyomachi, Wakayama, 640 - 8135 Japan E-mail: hori @ terra. zool. kyoto-u. ac. jp text Zootaxa 2019 2019-08-28 4661 2 271 308 journal article 21156 10.11646/zootaxa.4661.2.3 6c12aca1-7350-4265-9ded-6d415152f181 1175-5326 3998528 95722A8A-E02C-47E5-AA9B-777B6CA226E6 Cosmodela aurulenta ( Fabricius, 1801 ) ( Figs 8C , 13B ) Cicindela aurulenta Fabricius, 1801:239 . Cosmodela aurulenta aurulenta : Rivalier 1961:128 . Cosmodela aurulenta : Lopez-Lopez et al. 2015:8 . Specimens examined. (N = 4) [ Malaysia ] 2♂♂ ( YFJ ), Is. Penang , Penang National Park , 2. III. 2012 , Y. Fukuda leg ; [ Indonesia ] 2♂♂ ( EUMJ ), Yogyakarta , Mt. Merapi , Kaliurang , Java Is., 1. III. 2013 , R. Ogawa leg. Description . Internal sac everted widely and dorsally in lateral view. DA largest in bladders; B, VLL and A the second largest; DLL and VA the third largest; BLR, VLR, AL and VB smallest in bladders. A cylindrically everted apically. AL and VB slightly bulging at base of A, vested with fine spines, which are forming oblong spots. DA well swollen basally. DLL roundly swollen, vested numerously with spines, which are weakly chitinous. VA triangular swollen at base of DA, everted dorso-apically, tapered. BLR roundly swollen left ventrally, vested numerously with well sclerotized spines, which are forming an oblong spot. VLR swollen left laterally at base of VA. VLL oval and widely swollen right laterally, with well sclerotized spines, which are forming a narrow band. B oval and widely swollen right laterally, with well sclerotized spines, which are forming an oblong spot. Flagellum very long, curved and formed S-shaped. Medial tooth short, extending apically. Remarks. This species mostly shares the characteristics of the position of each bladder with C. batesi but is distinguished from the latter by VA and A not long.