Species of Staurotheca Allman, 1888 (Cnidaria: Hydrozoa: Sertulariidae) from US Antarctic expeditions, with the description of three new species
Author
Cantero, A. L. Peña
Author
Vervoort, W.
text
Journal of Natural History
2003
2003-11-30
37
22
2653
2722
http://dx.doi.org/10.1080/00222930210155701
journal article
10.1080/00222930210155701
1464-5262
Staurotheca profunda
sp. nov.
(figure 16;
table 5
)
Material examined. 22
/
1592
,
one fragment
ca
30 mm
long (
holotype
,
USNM 1003273
)
.
Description.
Material consisting of a single fragment
ca
30 mm
long, polysiphonic over almost its entire length. Main stem giving rise to three branches on the same side and another branch on the opposite side; branching approximately in one plane.
Hydrothecae alternately arranged in one plane (figure 16A, B), forming two longitudinal rows. Hydrothecae (figure 16A–C) only slightly immersed into the branches, strongly curved outwards. Adcauline hydrothecal wall free for approximately half of its length. Free adcauline wall straight or slightly convex. Abcauline hydrothecal wall concave. Aperture circular; rim even and usually provided with numerous short renovations, tilted downwards, forming an acute angle with long axis of branch. Hydrotheca lacking mushroom-shaped diaphragm.
Gonothecae not found.
Remarks.
In spite of the scarcity of available material, there is enough evidence to conclude that we are dealing with a new species.
Staurotheca profunda
sp. nov.
is characterized by the arrangement of the hydrothecae, in alternate pairs (cf.
table 6
), and by the shape of the hydrothecae that have a dish-shaped operculum, strongly pointing towards the inclusion of this species in the genus
Staurotheca
.
The alternate arrangement of the hydrothecae in two longitudinal rows is unusual in the genus (cf.
table 6
). Only
S. abyssalis
sp. nov.
has the same disposition as a general feature, although
S. australis
,
S. stolonifera
and
S. vervoorti
also have hydrothecae arranged in two longitudinal rows, usually in (sub)opposite pairs (in
S. australis
and
S. stolonifera
also in an alternate pattern).
Staurotheca amphorophora
may occasionally have two longitudinal rows of hydrothecae arranged in (sub)opposite pairs.
Staurotheca profunda
sp. nov.
differs from all those species by the shape of the hydrotheca. In
Staurotheca abyssalis
sp. nov.
the hydrotheca is adnate to the branch for
ca
one-quarter of its adcauline length. In
S. vervoorti
, by contrast, the hydrotheca is adnate for its full adcauline length. In
S. stolonifera
, the hydrotheca is free for
ca
one-third to one-quarter of its adcauline length. Finally, in
S. australis
the hydrotheca is different and it is adnate to the branch for a much longer part of the adcauline wall.
F. 16.
Staurotheca profunda
sp. nov.
(A) Part of branch showing alternate arrangement of hydrothecae; (B–C) several hydrothecae (note presence of operculum). (All drawings, Stn 22/1592.) Scale bar: 1 mm (A), 400 Mm (B–C).
Ecology and distribution.
Staurotheca profunda
sp. nov.
has been collected from depths between 1647 and
2044 m
at Burdwood Bank (Scotia Sea).
Etymology.
The specific name
profunda
refers to the deep waters from which it originates.
Table 5. Measurements of
Staurotheca profunda
sp. nov.
(in Mm).
Hydrothecae
Length of abcauline wall 900–950 Length of free adcauline wall 530–650 Length of adnate adcauline wall 650–740 Length of adcauline wall 1180–1310 Diameter at rim 320–370