Revision Of The Nearctic Species Of The Genus Amiota Loew (Diptera: Drosophilidae) Author Jones, Lance E. Author Grimaldi, David A. text Bulletin of the American Museum of Natural History 2022 2022-09-15 2022 458 1 181 https://bioone.org/journals/bulletin-of-the-american-museum-of-natural-history/volume-458/issue-1/0003-0090.458.1.1/Revision-of-the-Nearctic-Species-of-the-Genus-Amiota-Loew/10.1206/0003-0090.458.1.1.full journal article 10.1206/0003-0090.458.1.1 0003-0090 7400026 Amiota amputata , sp. nov. Figures 51A–B , 53A , 54 , 60D , 91A DIAGNOSIS: Large fly (ThL 1.51–1.71 mm ), dark brown, legs yellow; outer paraphysis laterally flattened, with hook on distal end, 2 short, blunt spines along middle portion of dorsal margin, base minutely serrate; inner paraphysis lost. DESCRIPTION: Large fly (ThL 1.51–1.71 mm ), dark brown, legs yellow. Frons light brown on ventral half, dorsal half of head darker. Facial marking large (FML/FMW 0.29–0.38). Cheek deep (EL/CW 7–13.66), posterior milky white. Marking under wing base prominent, large, milky white, margins clearly defined. Haltere yellow. Tergites 1 and 2 lightly colored. Arista: Short, plumose; longest branches D1 and D2; A.R. 0.28; 3 long dorsal, 2 short ventral branches; branch D3 pointed mediad; arista trunk with short-medium length microtrichia along most of its length. Male genitalia: Epandrium separated at midline with a large gap, margins distinct from surrounding membrane; long lobe ventrally, densely haired, single line of setae ascending dorsally; anterolateral projection small, oriented anteriorly. Cercus semicircular, margins entirely distinct from surrounding membrane. Surstylus approximately rectangular, ventral half covered in scattered setulae; 12 short, peglike prensisetae, apices blunt; small fingerlike lateral lobe, not extending beyond prensisetae. Subepandrial sclerite C-shaped in lateral view, with appendage heart shaped in posterior view, invaginated medially. Outer paraphysis laterally flattened with hook on distal end; 2 short, blunt spines along middle portion in lateral view; small proximal “wing” at base, minutely serrate. Inner paraphysis lost. Aedeagal apodeme as wide as long; curved 90° in lateral view, curved segment forming deep pockets; distal end slightly flared with a deep central emargination. Hypandrium thin, apex with gap; triangular in anterior, accommodating paraphysis; lateral arm with prominent gonopod, triangular, ventrally pointed. Ejaculatory apodeme long, 0.8× length of epandrium. Head and thorax measurements: (n = 5; Am 4, 457, 753, 1483,×1504) FL/FW 0.73 (0.66–0.81), EL/EW 1.30 (1.21–1.43), EL/CW 9.34 (7–13.66), FML/FMW 0.34 (0.29–0.38), PR / RR 0.60 (0.46–0.77), ThL 1.63 ( 1.51–1.71 mm ). TYPE MATERIAL : Holotype : male: Oak Cr. Canyon, Ariz., [ 34.912694 , -111.726958 ], VII-9- 41, R .H. Beamer, Am 457, [glued to paper point, dissected]. Deposited in the American Museum of Natural History ( AMNH ). Paratypes : 14 mi. SW. El Salto, Dgo. [ Durango ] MEX , [ 23.786449 , -105.597725 ], 8000′, June “26” 1964, J.F. McAlpine, 4♂ (Am 1448*, 1482*, 1483*, 1556*, CNC ) OTHER MATERIAL EXAMINED : Mexico : Chiapas : 5 mi NE San Cristobal , 1969-05-21 , leg. B. V . Peterson , 1♂ ( Am 1504*, CNC ) ; 5 mi. W San Cristobal , 7500′, 1969-05-06 , leg. H.J. Teskey , 1♂ ( Am 1427*, CNC ) . Durango : 24 mi W La Ciudad , 7000′, 1964-07-11 , leg. J.F. McAlpine , 1♂ ( Am 1553*, CNC ) . Puebla : 9 mi SE Huachinango , 1952-06, leg. M. Wasserman and W.B. Heed , 2260.13, 1♂ ( Am 4*, AMNH ) . USA : Arizona : Cochise Co. , Ramsay Canyon , Mile High Trail. Near Sierra Vista, 1984-07-24 , leg. L.B. Carlson , sweep, 1♂ ( Am 753*, DEBU ) ; Oak Creek Canyon, 1941-07-09 , leg. R . H. Beamer , 1♂ ( Am 457*, SEMC ) . FIG. 47. Heads and lateral views, A. subtusradiata species group. A–B. A. byersi , sp. nov. (Am 295, paratype, lateral image flipped). C–D. A. tibialis , sp. nov. (Am 495). ETYMOLOGY: From the Latin amputata , feminine past participle, for “pruned” or “amputated,” in reference to the stublike nature of the prongs on the outer paraphyses, which are much longer in other closely related species. DISTRIBUTION: This species has a large distribution from Chiapas State in Mexico north through central Arizona . COMMENTS: The paratype described in Máca (2003) as an “aberrant individual” from the type series of A. nebojsa Máca belongs is Amiota amputata . See Comments under that species.