Review of the tribe Smicromyrmini Bischoff, 1920 (Hymenoptera: Mutillidae) Author Lelej, Arkady S. 0000-0001-7501-0981 Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far East Branch of the Russian Academy of Sciences, Vladivostok, 690022, Russia. lelej @ biosoil. ru; https: // orcid. org / 0000 - 0001 - 7501 - 0981 lelej@biosoil.ru Author Williams, Kevin A. 0000-0002-6073-1070 Plant Pest Diagnostic Center, California Department of Food & Agriculture, 3294 Meadowview Road, Sacramento, CA 95832, USA. Kevin. Williams @ cdfa. ca. gov; https: // orcid. org / 0000 - 0002 - 6073 - 1070 illiams@cdfa.ca.gov text Zootaxa 2023 2023-01-27 5231 2 101 140 http://dx.doi.org/10.11646/zootaxa.5231.2.1 journal article 10.11646/zootaxa.5231.2.1 b5bfe9be-83bc-4ee5-b9e9-b5ea51dda9bc 1175-5326 7575308 EBD53535-397E-4894-B0A4-9AC3CC38DF3C 5. Dentilla Lelej in Lelej & Kabakov, 1980 ( Figs 11–16 , 111 ) Dentilla Lelej in Lelej & Kabakov, 1980: 195 , ♁, ♀ (as subgenus of Smicromyrme Thomson ); Lelej 1985: 190 , ♁, ♀ ; Lelej & Brothers 2008: 19 ; Brothers & Lelej 2017: 95 , ♁, ♀ ; Pagliano et al. 2020: 170 . Type species . Mutilla erronea André, 1900 [1902], ♁, by original designation (junior subjective synonym of Mutilla curtiventris André, 1901 according to Pagliano & Strumia (2007: 69) , resurrected to valid species by Lelej & Yildirim (2009: 15) . FIGURES 7–16. 7–10. Ctenoceraea pectinella Nonveiller, 1993 , ♁, holotype (7. Antenna, 8. Forewing, 9. Genitalia, dorsal view, 10. Genitalia, lateral view); 11–12. Dentilla rasnitsyni Lelej in Lelej & van Harten, 2011 , ♁, paratype (11. Lateral habitus, 12. Head, face view); 13–16. D. curtiventris (André, 1901) : 13. Dorsal habitus, ♀, 14. Lateral habitus, ♀, 15. Lateral habitus, ♁, 16. Head, face view. (Photographs by K. Williams, 7–10 from Nonveiller 1993 ). Diagnosis . MALE. Inner eye margin with weak notch. Mandible strongly widened apically, quadridentate, rarely tridenrtate, beneath with large subbasal tooth; preapical inner teeth equal to apical one or larger than it. Clypeus concave, anterior border usually with protruding medial part. Stigmatic cell 1.4 × distance between base of stigmatic cell and origin of base RS on Sc . T2 with long lateral felt lines, S2 with short (reduced) lateral felt lines. FEMALE. Head not widened posterad, posteriorly rounded. Pronotum with protruding humeral part, distinctly wider than propodeum. Scutellar scale more or less developed. T2 with one or three basal spots of pale setae, rarely with basal band of pale setae. Pygidial plate widely triangular, carinated laterally, surface with divergent striae. Diversity and distribution . Twenty-three species (13 based on males only, nine on females only, and one known from both sexes) are recognized predominantly from the Palaearctic Region ; there are six Afrotropical species (two of these penetrate from the Palaearctic); and Dentilla kompantsevi Lelej, 1995 and D. malinka ( Nurse, 1903 ) are Oriental . Remarks . All but three of the species are dull brown in color and apparently nocturnal in their behavior. These nocturnal species are somewhat commonly encountered in arid habitats from North Africa east to India ( Lelej 2002 , 2005 ). The more colorful diurnal species occur mainly in the Mediterranean ( Lelej, 2002 ). Lelej’s (1985) key includes about half of the currently recognized species.