Testing the phylogenetic utility of morphological character systems, with a revision of Creophilus Leach (Coleoptera: Staphylinidae) Author Clarke, Dave J. text Zoological Journal of the Linnean Society 2011 Zool. J. Linn. Soc. 2011-10-25 163 3 723 812 http://dx.doi.org/10.1111/j.1096-3642.2011.00725.x journal article 10.1111/j.1096-3642.2011.00725.x 0024-4082 5441985 FBFE9195-BE04-4AFE-9417-6E38BCE6AB84 CREOPHILUS MAXILLOSUS SPECIES- GROUP This species group was originally proposed by Newton (1985) for the Holarctic and Oriental species. With recognition of three new synonyms and description of one new species, five species are included here. Diagnosis: Head black or brownish-black; antennomere 2 subequal in length to 3 or shorter; antennomere 11 with apical pairs of setae situated on either side of antennal axis (e.g. Fig. 13F ); right mandible with T3 directly ventral to T2 and mandibular blade, mostly (in C. incanus entirely) hidden from dorsal view beneath mandibular blade ( Fig. 13A ); anterior pronotal angles produced laterally ( Figs 2G , 13E ); abdominal tergites III–V without secondary transverse basal ridge; apical fringe of tergites III–VI with at least paired groups of white or golden setae medially, rest of body with variably black, brown, whitish-grey, or golden-brown vestiture ( Fig. 1F–J ); median lobe of aedeagus completely divided ventrally by membranous strip ( Fig. 3C , arrow). Large males with: apical portion of mandibles exceeding 0.9 times length of basal portion (ML1/ML2 range of all males = 0.60–1.09); without ventrolateral carina of head; pronotum widest at anterior angles; base of profemur with spine-like apophysis ( Fig. 9D , arrow); superior pronotal marginal line abruptly deflected beneath disc of pronotum at point of basolateral denticle ( Fig. 2G , bpd ).