An overview of the extant genera and subgenera of the order Scolopendromorpha (Chilopoda): a new identification key and updated diagnoses
Author
Schileyko, Arkady A.
schileyko1965@gmail.com
Author
Vahtera, Varpu
varpu.vahtera@gmail.com
Author
Edgecombe, Gregory D.
0000-0002-9591-8011
schileyko1965@gmail.com
text
Zootaxa
2020
2020-08-10
4825
1
1
64
journal article
8703
10.11646/zootaxa.4825.1.1
5ab5f5c8-481e-4d1a-8643-21e72c367278
1175-5326
4402145
F230F199-1C94-4E2E-9CE4-5F56212C015F
Subfamily
Newportiinae Pocock, 1896
Synonyms.
Ectonocryptopinae
Shelley & Mercurio, 2005
Diagnosis.
Antenna with normal trichoid sensilla, lacking collared sensilla (
Fig. 23
). Anterior margin of forcipular coxosternite without well-developed tooth plates, either smooth (
Figs 17
,
21
) or with projections in
Newportia
(
Tidops
) Chamberlin, 1915
(
Fig. 22
). Forcipular trochantero-prefemur without well-developed process (with pointed minute tubercle in some species); tarsungula of variable length (
Figs 21, 22
). LBS 7 with (in most species) or without spiracles. Coxopleuron with conical process of variable length. Ultimate legs “lasso-shaped” sensu
Schileyko (2009)
(i.e. their tarsus 2 elongated and divided into numerous (4 to 40) variably-distinct secondary articles thus forming a kind of functional antenna,
Figs 19
,
20
) or of “ectono-type” (see
Schileyko 2009
), i.e. somewhat shortened and swollen, subclavate, with inflated distal podomeres (
Figs 14, 15
). Ultimate prefemur with 3–6 large spinous processes, femur with a few (0–3) smaller spinous processes. Ultimate pretarsus absent in most taxa of both genus-group and species (except for 3 or 4 species of
Newportia
(
Newportides
)
Chamberlin, 1912
, figs
29, 47 in
Chagas-Jr 2018).
Number of subtaxa.
1 genus, 5 subgenera (“Ca 60 species in two genera” in
Edgecombe & Bonato 2011: 405
).
Sexual dimorphism.
Unknown.
Range.
Neotropics: from Central Mexico to
Paraguay
, including Cocos Island and Caribbean Islands.
FIGURES 7–13.
Scolopocryptops melanostoma
Newport, 1845
; Ad. Rc 7172
7
LBS 23 and ultimate legs, laterally; Ad. Rc 7503
8
anterior margin of forcipular coxosternite + maxillae 2, ventrally;
Scolopocryptops ferrugineus
(L., 1767); Peru specimen from Markus Koch collection
9
forcipular segment ventrally;
Scolopocryptops miersii
Newport, 1845
; BMNH, BM 1889.4.13.3–5
10
apical part of maxilla 2 laterally;
Kethops utahensis
(Chamberlin, 1909)
11
figures 4–6 of
Chamberlin (1912)
;
Thalkethops grallatrix
Crabill, 1960
12
figure 15 of
Crabill (1960)
;
Kethops utahensis
(Chamberlin, 1909)
; USNM Mill Creek (Utah) specimen
13
apical part of maxilla 2 ventro-laterally; (
20
)—leg 20, (
am
)—anterior margin of forcipular coxosternite, (
cp
)—coxopleural process, (
cst
)—forcipular coxosternite, (
dms
)—dorso-medial spinous process of ultimate prefemur, (
p2
)— pretarsus of maxilla 2, (
pa
)—short projection of anterior margin of forcipular coxosternite, (
pf
)—prefemur, (
pla
)—pretarsal lamellae, (
pt
)—process of forcipular trochantero-prefemur, (
rl
)—right[?] ultimate leg, (
S
)—? sternite, (
st
)—saw teeth-like small spinous processes, (
t1
)—ultimate tarsus 1, (
t2
)—ultimate tarsus 2, (
ti
)— ultimate tibia, (
us
)—ultimate LBS, (
ust
)—ultimate sternite, (
vsp
)—ventral spinous process of ultimate prefemur.
Remarks.
Treated as a subfamily in
Edgecombe & Bonato (2011: 405)
,
Vahtera
et al
. (2012a: 9
,
2013: 578
). As a result of work of
Vahtera
et al.
(2013)
this subfamily became monotypic because the former genera
Tidops
,
Ectonocryptops
Crabill, 1977
and
Ectonocryptoides
should belong to a single genus
Newportia
Gervais, 1847
(see below).