Deep-sea Ophiuroidea (Echinodermata) collected during the TALUD cruises in western Mexico
Author
Granja-Fernández, Rebeca
0000-0001-7119-0567
beckygranja@gmail.com
Author
Hendrickx, Michel E.
0000-0001-9187-6080
michel@ola.icmyl.unam.mx
Author
Rangel-Solís, Pedro Diego
0009-0008-8478-0679
olis@gmail.com
Author
López-Pérez, Andrés
0000-0001-7119-0567
beckygranja@gmail.com
text
Zootaxa
2023
2023-04-03
5259
1
1
71
http://dx.doi.org/10.11646/zootaxa.5259.1.1
journal article
56236
10.11646/zootaxa.5259.1.1
e2c999aa-4046-4fc1-a351-158efbf234cb
1175-5326
7794984
4306F52E-FD24-45B0-B307-66B71173D805
Ophiopholis bakeri
McClendon, 1909
Fig. 18G‒L
Ophiopholis bakeri
McClendon, 1909: 41
, pl. 5, figs. 26‒27.— H.L.
Clark 1911: 121‒123
, fig. 46.—
Ziesenhenne 1937: 224
.—
Boolotian & Leighton 1966: 7‒8
, fig. 28.—
Kyte 1969a: 1729
, 1732, 1738.—
Hendler 1996: 157‒159
, fig. 7.17.—
Lambert & Austin 2007: 110
, 113‒114, figs. 62‒63.
See
Hendler (1996)
for other synonymous records.
Material examined
. 150 individuals at three stations.
TALUD XIV
, Sta. 4, 2 ind. (ICML-EMU-11043-B); Sta. 10, 10 ind. (ICML-EMU-13003); Sta. 15, 1 ind. (ICML-EMU-11043-A); Sta. 20, 22 ind. (ICML-EMU-11042-B), 87 ind. (ICML-EMU-11041), and 10 ind. (ICML-EMU-11042-A).
TALUD XV
, Sta. 23, 18 ind. (ICML-EMU-11043-C).
Comparative material.
MCZ
OPH-7061
,
2
ind (
Supplementary
file 2). Material identified by McClendon. The lot
MCZ
OPH-7061
deposited at the
MCZ
has a label with the legend: “Univ. Calif. Dep. Zool.
Type
?”. However, it does not have any additional information (e.g., locality, collection date) or any other note suggesting that it is, in fact, the
type
material of
O
.
bakeri
. McClendon, who originally described the species (
McClendon 1909
), is actually the person who identified lot
MCZ
OPH-7061
(pers. comm. Penny Benson 2016). According to
McClendon (1909)
, the
type
material of
O
.
jolliense
was deposited at the University of
California
Museum but despite an exhaustive search for this material, we did not locate it in this museum (pers. comm.
Erica Clites
&
Carole Hickman
2018) or in any other institution (i.e.,
LACM
,
MCZ
,
NHMD
,
USNM
; pers. comm.
Penny Benson
2016,
Rich Mooi
2018). Lot
MCZ
OPH-7061
might have been sent to
MCZ
as a loan or a gift.
Description
(ICML-EMU-11041). DD =
8 mm
. Disc round, weakly indented interradially. Dorsal disc covered by scales and short multifid spines, marginal and central spines elongated. Primary plates not evident. RS scalene triangular, covered by multifid spines (often shorter than on other parts of the disc), separated by scales and multifid spines, except distally (
Fig. 18G
). Ventral interradii covered by elongated multifid spines (
Fig. 18H
). OSh broader than long, diamond-shaped. Madreporite larger and rounder than OSh.AdSh curved, separated in front of OSh. Jaws bearing numerous oral papillae; one AdShSp rounded; one 2AdShSp oblong, the longest; LOPa 1‒2 oblong; TPas numerous, clustering, short. vT rectangular (
Fig. 18I
). Arms slender. DAP as broad as long, rounded, surrounded by rounded symmetrical plates, contiguous (
Fig. 18J
). VAP longer than broad, pentagonal, separated.LAP with up to 4‒5 ArSp, blunt tip, serrated, elongated (approximately two arm segments in length), some first dorsalmost rudimentary, second dorsalmost the longest, ventralmost the shortest, distalmost arm spines hooked, hyaline. Tentacle pores with one oval TSc (
Fig. 18K
). Color pattern dorsally and ventrally beige-whitish (ethanol preservation) (
Fig. 18G‒L
).
FIGURE 18.
Histampica
cf.
duplicata
(
Lyman, 1875
)
(ICML-EMU-12996, DD = 5.4 mm). A, Dorsal disc; B, Ventral disc; C, Mouth; D, Dorsal arm; E, Ventral arm; F, Dorsal view.
Ophiopholis bakeri
McClendon, 1909
(ICML-EMU-11041, DD = 8 mm). G, Dorsal disc; H, Ventral disc; I, Mouth; J, Dorsal arm; K, Ventral arm; L, Dorsal view.
Habitat and distribution.
British Columbia
,
Canada
,
Washington
,
Oregon
, and
California
,
USA
, and northern Mexican Pacific; 18‒
1,204 m
depth, coral, sponges, rocky, sandy, and muddy substrates (
McClendon 1909
; H.L.
Clark 1911
;
Hendler 1996
;
Lambert & Austin 2007
;
Granja-Fernández
et al.
2015
). The material examined was collected off western Baja
California
Sur and in the northern Gulf of
California
;
325‒871 m
depth, associated with sponges.
Remarks.
Many of the ethanol preserved specimens had dorsally and ventrally beige-whitish coloration. Nevertheless, we also observed that some preserved specimens displayed a rosaceous or brown coloration. This species is known to display a wide variety of colors: pink, red, gray, and yellowish-white (
Hendler 1996
;
Lambert & Austin 2007
). Moreover, multifid spines in the center of the disc can be present or not, and some specimens can have up to six arm spines. The
Ophiopholis
species
distributed in the eastern tropical Pacific (northern
Mexico
) are
Ophiopholis bakeri
and
Ophiopholis longispina
, which differentiate by: 1) dorsal disc covered with scales and short multifid spines in
O
.
bakeri
vs. covered with scales and long spines in
O
.
longispina
, 2) RS covered with multifid spines in
O
.
bakeri
vs. RS without spines in
O
.
longispina
, and 3) LAP with up to 4‒5 ArSp, some first dorsalmost rudimentary, second dorsalmost the longest in
O
.
bakeri
vs. up to 6‒7 ArSp, first dorsalmost the longest in
O
.
longispina
. On the other hand,
Ophiopholis aculeata
(
Linnaeus, 1767
)
and
Ophiopholis kennerlyi
Lyman, 1860
have been reported from the North Pacific and differ from
O
.
bakeri
by their granule-covered dorsal disc and their angular accessory arm plates (
Lambert & Austin 2007
).