Revision of the New Caledonian endemic genus Gracilipsodes (Trichoptera: Leptoceridae: Grumichellini)
Author
Malm, Tobias
Author
Johanson, Kjell A.
text
Zoological Journal of the Linnean Society
2008
2008-07-31
153
3
425
452
https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2008.00403.x
journal article
10.1111/j.1096-3642.2008.00403.x
0024-4082
5446337
GRACILIPSODES AURORUS
SP. NOV.
(
FIGS 4
,
13
)
Diagnosis:
This species is distinguished from
G. lanceolatus
sp. nov.
,
G. grandis
sp. nov.
,
G. psocopterus
, and
G. similis
by the tibial spur formula 2, 2, 2 and the presence of a pair of pre-apical lateral processes of tergum X, originating from the apical half of the segment. It is separated from
G. aureus
sp. nov.
and
G. robustus
sp. nov.
in the genitalia, by the longer pre-apical lateral processes of tergum X, and in the forewing, by a longer crossvein m–cu, oriented at a sharp angle to M. It is separated from
G. koghiensis
sp. nov.
and
G. aoupiniensis
sp. nov.
by having a darker body colour, by the notched tergum X apex, and by the smaller number of both hind tibial spines and mesal spines on the apical part of the inferior appendage.
Description, male:
Body dark greyish brown. Forewing transparent grey with pale spots. Tibial spur formula 2, 2, 2. Hind tibiae each with 15–20 small blackish spines (
N
= 27). Forewing (
Fig. 4A
): length 7.0–
7.6 mm
(
N
= 27), apically rounded; forks 1 and 5 present; crossvein m–cu nearly as long as crossvein r–m, and sharply angled to M. Hindwing (
Fig. 4B
): length
5.3–5.7 mm
(
N
= 27), apex nearly acute; forks 1, 3, and 5 present; crossvein r–m convex; costa with 16 hamuli.
Genitalia:
Segment IX annular, laterally setose, narrowest at bases of superior appendages; in lateral view tergite IX wider than sternite IX (
Fig. 4C
), central part of tergite IX posterior margin produced posteroventrally (
Fig. 4C, D
), smooth, except for a few long setae in some individuals. Superior appendages setose, flattened dorsoventrally, as long as or longer than tergum X, and apices narrowly rounded in dorsal view (
Fig. 4D
). Tergum X membraneous; median process entire, wide at basis, tapering to rounded apex, in ventral view with straight dorsal margin (
Fig. 4C
), and apex with a wide V-shaped notch in dorsal and ventral views (
Fig. 4D, E
); pre-apical lateral processes about as long as median process, produced at apical half, sclerotized, digitate, directed posterad, and slightly curved ventrad (
Fig. 4C, E
). Inferior appendages each bipartite, and setose; basal part basally slender in lateral view (
Fig. 4A
), wide in ventral view (
Fig. 4E
), mesally concave and produced posteromesally into broad, apically rounded lip, well-sclerotized along margin, and bearing small dorsal spines (
Fig. 4C, E
); apical part of each inferior appendage digitate, curving mesally; in lateral view, straight, parallel-sided, apex rounded, with 25–30 mesal spines (
Fig. 4C
); mesal process with nine strong spines (
Fig. 4D
). Harpago: small, sclerotized (
Fig. 4C
). Phallic apparatus simple (
Fig. 4F, G
); phallicata basally tubular, fused with phallobase, and apex truncate; bent about 45° ventrally at distal half; apically with pair of sclerotized, broad lateral processes, ventrally directed, tapering distally (
Fig. 4F
); phallotremal sclerite U-shaped (
Fig. 4G
).
Figure 3.
Gracilipsodes aureus
sp. nov.
A, forewing. B, hindwing. C, genitalia, lateral. D, genitalia, dorsal. E, genitalia, ventral. F, phallus, lateral. G, phallus, ventral.
Figure 4.
Gracilipsodes aurorus
sp. nov.
A, forewing. B, hindwing. C, genitalia, lateral. D, genitalia, dorsal. E, genitalia, ventral. F, phallus, lateral. G, phallus, ventral.
Holotype
(male):
New Caledonia
,
Province Nord
,
Mt Panié
, stream at camp,
20.58167°S
,
164.76472°E
,
1311 m
a.s.l.
,
9 December 2003
–
2 January 2004
,
Malaise trap
, loc#073 (
K. A. Johanson
).
Paratypes
:
12 males
, same data as holotype. Fourteen males, ditto, except
20.58139°S
,
164.76444°E
,
1310 m
a.s.l.
, loc#074
.
Distribution:
Province Nord
, Mt Panié.
Etymology:
aurorus
, from Latin,
aurora
, dawn, daybreak; referring to the dark, sclerotized basal posteromesal lip of the inferior appendage, which contrasts with the paler basal part.