Lepocranus and Valalyllum gen. nov. (Orthoptera, Tetrigidae, Cladonotinae), endangered Malagasy dead-leaf-like grasshoppers Author Deranja, Maks https://orcid.org/0000-0002-5710-1916 University of Zagreb, Faculty of Science, Department of Biology, Division of Zoology, Evolution Lab, Rooseveltov trg 6, HR- 10000 Zagreb, Croatia maks.deranja@gmail.com Author Kasalo, Niko https://orcid.org/0000-0002-3139-6349 University of Zagreb, Faculty of Science, Department of Biology, Division of Zoology, Evolution Lab, Rooseveltov trg 6, HR- 10000 Zagreb, Croatia niko.kasalo5@gmail.com Author Adzic, Karmela https://orcid.org/0000-0001-6223-4759 University of Zagreb, Faculty of Science, Department of Biology, Division of Zoology, Evolution Lab, Rooseveltov trg 6, HR- 10000 Zagreb, Croatia Author Franjevic, Damjan https://orcid.org/0000-0001-6989-4426 University of Zagreb, Faculty of Science, Department of Biology, Division of Zoology, Evolution Lab, Rooseveltov trg 6, HR- 10000 Zagreb, Croatia Author Skejo, Josip https://orcid.org/0000-0002-2554-4499 University of Zagreb, Faculty of Science, Department of Biology, Division of Zoology, Evolution Lab, Rooseveltov trg 6, HR- 10000 Zagreb, Croatia text ZooKeys 2022 2022-07-01 1109 1 15 http://dx.doi.org/10.3897/zookeys.1109.85565 journal article http://dx.doi.org/10.3897/zookeys.1109.85565 1313-2970-1109-1 FC606834C2594254B8CA9C5A6007246B 8895C118E1765B7A87C2D49A619D0EA4 Subfamily Cladonotinae Composition and distribution. This diverse and polyphyletic family includes altogether 314 species assigned to 74 genera ( Cigliano et al. 2022 ). Most genera require revision and are not adequately classified in the Tetrigidae system ( Zhang et al. 2020 ). Within the subfamily Cladonotinae , several monophyletic tribes exist, such as Caribbean Choriphyllini ( Choriphyllum Serville, 1838 and Phyllotettix Hancock, 1902), African Xerophyllini (including leaf-like Acmophyllum Karsch, 1890, Seyidotettix Rehn, 1938, Trypophyllum Karsch, 1890, Xerophyllum Fairmaire, 1846), and Asian Cladonotini ( Hymenotes Westwood, 1837, Holoarcus Hancock, 1909, Dolatettix Hancock, 1907). Diversity in Madagascar. Altogether, 18 Tetrigidae species of Madagascar are assigned to the subfamily Cladonotinae : two members of Valalyllini trib. nov. ( L. fuscus and V. folium ); 13 species of the genus Thymochares , which is currently not assigned to any of the tribes, and it is questionable whether it represents a genus of Cladonotinae ; Microthymochares pullus , an endemic genus and species with the same taxonomic problem as Thymochares ; Epitettix spheniscus also with the same problematic assignment; and Xerophyllini and its member Morphopoides madagascariensis ( Cigliano et al. 2022 ).