A revision of the Cossulinae of Costa Rica and cladistic analysis of the world species (Lepidoptera: Cossidae) Author Davis, Steve R. Author Gentili-Poole, Patricia Author Mitter, Charles text Zoological Journal of the Linnean Society 2008 2008-10-31 154 2 222 277 https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2008.00406.x journal article 10.1111/j.1096-3642.2008.00406.x 0024-4082 5446371 BIOCELLATA BIFIDA SP. NOV. ( FIGS 22 , 55, 56 , 90 , 136 ) Male ( Fig. 22 ): Forewing length: 15–17 mm . Head: Antenna with shaft and rami creamy white; labial palpus brown dorsally, creamy white with some light brown ventrally; frons creamy white; vertex creamy white, irrorated with brownish grey. Thorax: Patagia creamy white, heavily irrorated with brownish grey; dorsum and tegula with scales creamy white and creamy white basally, greyish brown apically; pleura cream; venter mostly brown with some cream. Legs with femur and tibia cream, irrorated with brown; fore tarsi cream, light brown, and brown; mid and hind tarsi brown, banded creamy brown distally on each tarsomere. Forewing creamy white with numerous grey transverse striae throughout wing dorsally; the two most distinct and largest of these striae located basally and medially along wing; subcostal region creamy white and mostly lacking striae; numerous short, transverse, grey and dark brown striae along costal margin; two dark brown to fuscous spots subterminally; fringe banded light creamy gold and brownish grey; forewing mostly light brown ventrally with some cream; numerous short, transverse, dark brown striae along costal margin. Hindwing creamy white dorsally; some creamy light brown along apical margin; fringe can be slightly banded grey and creamy white, but mostly creamy white; hindwing mostly creamy white ventrally. Figures 95–100. Male genitalia, lateral view of genital capsule: 95, Spinulata quasivinnea ; 96, Simplicivalva ampliophilobia ; 97, Magulacra niveogrisea ; 98, Cossula arpi ; 99, Cossula buspina ; 100, Cossula coerulescens . Scale bar = 1.0 mm. Abdomen: Mostly cream and creamy white dorsally; cream and brown ventrally; A8 cream with slight light brown. Male genitalia ( Figs 55, 56 , 90 ): Valva subtrapezoidal, approximately 0.9¥ length of genital capsule; gnathos–uncus expanse approximately 0.5¥ length of genital capsule; saccular process absent, with sacculus heavily sclerotized and bearing flattened rugosities; juxta process small and slender; bridge of gnathos strongly bifurcate, with extending arms slightly upcurved and apex of each arm acute; each arm with a serrate ridge ventrally; uncus shallowly bifurcate, with apices acute. Aedoeagus with sclerotized plate formed mediad, where juxta attaches; rostellum long and slender, inwardly curving, with apex acuminate as viewed laterally; a patch of triangular spines on one side, along margin where vesica emerges from aedoeagus; rostellum becoming slightly more narrow than aedoeagus as viewed ventrally, with apex rounded. Female: Unknown. Distribution: Known only from Heredia and Limon Provinces, Costa Rica .