Review of the genus Hyrtanella (Ephemeroptera: Ephemerellidae) Author Jacobus, Luke M. Author Sartori, Michel text Zootaxa 2004 785 1 12 journal article 10.5281/zenodo.158566 ac5a8900-3ed1-4993-82a2-3de55f724002 1175­5326 158566 4D1175CD-8F0B-4969-9FDE-34C225D2E52C Hyrtanella Allen and Edmunds Diagnosis. Larvae are distinguished by having claws with basal denticles and a set of long subapical denticles; gills 3 operculate, with a median, transverse, weakened band; paired, blunt spines on abdominal terga 4–7; single, median spines on abdominal terga 8 and 9; and abdominal segments 8 and 9 reduced in width ( Allen & Edmunds 1976 ). Female adults are distinguished by having a reduced number of marginal intercalaries between CuA and IMP in the forewing (one intercalary between IMP and MP2, and no intercalaries between MP2 and CuA), a relatively small hindwing with few crossveins and a sharp costal projection, a constricted abdomen, a medial spine vestige on abdominal segment 8, and enlarged abdominal segments 8 and 9 ( Allen & Edmunds 1976 ; Allen 1980 ). Discussion. In the original description, Allen and Edmunds (1976) erroneously described gills present only on segments 3–6. As noted by McCafferty and Wang (2000) and Kluge (2004) , Hyrtanella larvae indeed possess gills on segments 3–7.