A sabre-tooth predator from the Neotropics: Cranial morphology of Anachlysictis gracilis Goin, 1997 (Metatheria, Thylacosmilidae), based on new specimens from La Venta (Middle Miocene, Colombia) Author Suarez, Catalina Current affilition: Instituto Argentino de Nivología, Glaciología y Ciencias Ambientales (IANIGLA), CCT-CONICET Mendoza, Av. Ruiz Leal s / n, Parque General San Martín, Mendoza, 5500 (Argentina) and Grupo de Paleontología Neotropical Tradicional y Molecular (PaleoNeo), Facultad de Ciencias Naturales, Universidad del Rosario, Bogotá (Colombia) and Smithsonian Tropical Research Institute, Box 0843 - 03092, Balboa-Ancón (Panama) catasuarezg @ gmail. com (corresponding author) catasuarezg@gmail.com Author Forasiepi, Analia M. Instituto Argentino de Nivología, Glaciología y Ciencias Ambientales (IANIGLA), CCT-CONICET Mendoza, Av. Ruiz Leal s / n, Parque General San Martín, Mendoza, 5500 (Argentina) and Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET) (Argentina) Author Babot, María Judith Unidad Ejecutora Lillo-CONICET, Fundación Miguel Lillo, Miguel Lillo 251, San Miguel de Tucumán, Tucumán, 4000 (Argentina) Author Shinmura, Tatsuya Ashoro Museum of Paleontology, 29 - 1 Konan, Ashoro, Ashoro District, Hokkaido 089 - 3727 (Japan) Author Luque, Javier Museum of Zoology, Department of Zoology, University of Cambridge, Downing Street, Cambridge CB 2 3 EJ (United Kingdom) and Smithsonian Tropical Research Institute, Box 0843 - 03092, Balboa-Ancón (Panama) Author Vanegas, Rubén D. Museo de Historia Natural La Tatacoa, Centro Poblado La Victoria, Villavieja, Huila Department (Colombia) Author Cadena, Edwin A. Grupo de Paleontología Neotropical Tradicional y Molecular (PaleoNeo), Facultad de Ciencias Naturales, Universidad del Rosario, Bogotá (Colombia) and Smithsonian Tropical Research Institute, Box 0843 - 03092, Balboa-Ancón (Panama) and Field Museum of Natural History, Chicago, IL 60605 (United States) Author Goin, Francisco J. Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET) (Argentina) and División Paleontología de Vertebrados, Museo de La Plata, Facultad de Ciencias Naturales y Museo, Universidad Nacional de La Plata, Paseo del Bosque s / n, La Plata, Buenos Aires, B 1900 FWA (Argentina) text Geodiversitas 2023 2023-10-12 45 18 497 572 https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2023v45a18.pdf journal article 10.5252/geodiversitas2023v45a18 1638-9395 8434228 urn:lsid:zoobank.org:pub:BB77B691-635A-4B92-9820-DBE74776B7E2 Patagosmilus goini Forasiepi & Carlini, 2010 MLP 07-VII-1-1 ( holotype ), most of the left side of the skull with upper dentition, right magnum, and proximal portion of ungual phalanx ( Río Negro Province , Argentina ; Collón Curá Formation, Colloncuran SALMA, middle Miocene) ; B:p2-154 ( MNHN-Bol : cast hosted in the MLP and observations from F. Goin), posterior portion of a skull, a fragment of left maxilla with the proximal portion of the canine, fragments of left and right maxillae with dentition, incomplete left and right mandibular rami with dentition (“ Bone wash ” locality, Río Rosario , Tarija Department , Bolivia ; un-named formation, Honda Group, Bolivia [not homologous to the Honda Group from Colombia (MacFadden & Wolff 1981)], Laventan SALMA, middle Miocene) . Additional information: MacFadden & Wolff (1981); Goin & Carlini (1993); Forasiepi & Carlini (2010). Comment: P.goini is currently known for the holotype and the only published specimen. However, another still-undescribed specimen from Quebrada Honda, Bolivia (middle Miocene) has been mentioned by Goin & Carlini (1993) as belonging to the same, or closely related species than the future holotypes of A. gracilis (named a few years later by Goin 1997) and P. goini (named some years later by Forasiepi & Carlini 2010). The specimen is identified with the field number “B: p2-154 ( July 1992 )” and was collected during the Duke University/Geobol Paleontology Expedition in 1992 (collectors: Richard Kay, Richard Madden, Carlos Villarroel, Federico Anaya, Marcelo Sánchez, Blythe Williams). The fossil stayed in the La Plata Museum (MLP) for study by Dr Francisco Goin's team. Still, that research could not be finished because the specimen was requested by the Natural History of Bolivia (MNHN-Bol), which was designed as its official repository until now. During the development of this research, we did not have access to the original material of this specimen, trying to establish communication with that museum several times over several years without success. We hope this communication is finally possible soon, so the study of this specimen can be finally completed. However, we had access to a cast hosted in the MLP and the notes of observations made by Dr Goin while he was studying the fossil. We used that material and information for comparisons and to complete some scorings in the matrix for P. goini , as we agree with the hypothesis of this specimen belonging to P. goini .